New Syrphidae (Diptera) of North-eastern North America

Biodiversity Data Journal 7: e36673 OO)
doi: 10.3897/BDJ.7.e€36673 open access
Taxonomic Paper

New Syrphidae (Diptera) of North-eastern North
America

Jeffrey H. Skevington*8, Andrew D. Young], Michelle M. Locke*, Kevin M. Moran*®

+ AAFC, Canadian National Collection of Insects, Arachnids and Nematodes, Ottawa, Canada
§ Carleton University, Ottawa, Canada
| California Department of Food and Agriculture, Sacramento, United States of America

Corresponding author: Jeffrey H. Skevington (jhskevington@gmail.com)
Academic editor: Torsten Dikow
Received: 31 May 2019 | Accepted: 09 Aug 2019 | Published: 03 Sep 2019

Citation: Skevington JH, Young AD, Locke MM, Moran KM (2019) New Syrphidae (Diptera) of North-eastern North
America. Biodiversity Data Journal 7: e36673. httos://doi.org/10.3897/BDJ.7.e36673

ZooBank: urn:lsid:zoobank.org:pub:823430AD-B648-414F-A8B2-4F1E5F1A086A

Abstract
Background

This paper describes 11 of 18 new species recognised in the recent book, "Field Guide to
the Flower Flies of Northeastern North America". Four species are omitted as they need to
be described in the context of a revision (three Cheilosia and a Palpada species) and three
other species (one Neoascia and two Xy/ota) will be described by F. Christian Thompson in
a planned publication. Six of the new species have been recognised for decades and were
treated by J. Richard Vockeroth in unpublished notes or by Thompson in his unpublished
but widely distributed "A conspectus of the flower flies (Diptera: Syrphidae) of the Nearctic
Region”. Five of the 11 species were discovered during the preparation of the Field Guide.
Eight of the 11 have DNA barcodes available that support the morphology.

New information

New species treated in this paper include: Anasimyia diffusa Locke, Skevington and
Vockeroth (Smooth-legged Swamp Fly), Anasimyia matutina Locke, Skevington and

This is an open access article distributed under the terms of the CCO Public Domain Dedication.

2 Skevington J et al

Vockeroth (Small-spotted Swamp Fly), Brachyopa caesariata Moran and Skevington
(Plain-winged Sapeater), Brachyopa cummingi Moran and Skevington (Somber Sapeater),
Hammerschmidtia sedmani Vockeroth, Moran and Skevington (Pale-bristled Logsitter),
Microdon (Microdon) scauros Skevington and Locke (Big-footed Ant Fly), Mixogaster fattigi
Locke, Skevington and Greene (Fattig's Ant Fly), Neoascia guttata Skevington and Moran
(Spotted Fen Fly), Orthonevra fee; Moran and Skevington (Fee's Mucksucker), Psilota
klymkoi Locke, Young and Skevington (Black Haireye) and Trichopsomyia_litoralis
Vockeroth and Young (Coastal Psyllid-killer). Common names follow the "Field Guide to the
Flower Flies of Northeastern North America" (Skevington et al. 2019).

Keywords

new species, taxonomy, flower flies, hover flies, Diptera, Syrphidae, Nearctic

Introduction

Flower flies (Syrphidae) are important pollinators, perhaps even outperforming native bees
in this critical agricultural ecosystem service (Rader et al. 2016, Skevington et al. 2019). In
addition to this important role, most immature Syrphinae species are predators of soft-
bodied insects such as aphids and scales, many of which are agricultural pests (Rojo et al.
2003, Tenhumberg and Poehling 1995). Many Eristalinae syrphid larvae, such as Eristalis
Latreille 1804 and Eristalinus Rondani 1845, perform critical roles in recycling human and
farm animal waste (Pérez-Bahon 2013). Other saprophagous species are found in old
growth forests where they specialise in rotholes or under the bark of large trees and are
important indicators of habitat health (e.g. Blera fallax Linnaeus 1758; Rotheray et al.
2012). Overall, syrphids have offered important insights into the health of our ecosystems
and declines in insect numbers (Hallmann et al. 2017, Sanchez-Bayo and Wyckhuys
2019). A few syrphid larvae are phytophagous (with some Cheilosia Meigen 1822 used in
biological control of plants (Grosskopf 2005)), while those of the subfamily Microdontinae
are specialised ant predators and even parasitoids (Pérez-Lachaud et al. 2014). In fact,
syrphid larvae are more varied and have more ecological roles than any other family of
flies, with the possible exception of Phoridae (McAlpine et al. 1981, McAlpine et al. 1987).
The family is species-rich, as well as ecologically diverse, with 828 valid species of
Nearctic Syrphidae currently recognised (Skevington 2019) and over 6200 species known
worldwide (Pape and Evenhuis 2019). Many syrphids are either perfect or imperfect
mimics of bees and wasps, adding to both their interest scientifically (e.g. Edmunds and
Reader 2014, Gilbert 2005, Penney et al. 2012, Penney et al. 2014) and to confusion by
non-specialists who think that all floral visitors are bees. Many ‘bees’ portrayed in
advertisements and even in the scientific literature are actually syrphids.

The popularity of Syrphidae, both as research subjects and as the subject of amateur
natural history pursuits, has been increasing in recent years and field guides are emerging
that capitalise on this interest. The first such guide was targeted on the United Kingdom

New Syrphidae (Diptera) of North-eastern North America 3

fauna and, although not comprehensive, provided an excellent entry into the taxonomy and
identification of the group (Ball and Morris 2015). The year 2019 has seen the publication
of two comprehensive guides to syrphids. The first, by Bot and Van de Meutter (2019),
treated all 384 species that are found, or are expected to be found, in The Netherlands and
Belgium. The second, by Skevington et al. (2019) (hereafter, the “Field Guide”), treated
413 species found in north-eastern North America. This book includes 18 undescribed
species of Syrphidae that are illustrated but not named. This paper names 11 of these 18
new taxa. Seven species are omitted (three Cheilosia, a Neoascia Williston 1887 species,
a Palpada Macquart 1834 species and two Xylota Meigen 1822 species). Chei/osia is one
of the most diverse Nearctic syrphid genera and includes 119 names (Pape and Evenhuis
2019). Moran reworked the eastern Nearctic species for inclusion in the Field Guide but did
not study all related type material. Western Nearctic and Palaearctic species need to be
included in a complete revision before we can be absolutely sure that morphospecies
included in the field guide are not valid but are currently undiagnosable species. Palpada is
not as complicated, but all types previously assigned as synonyms to P. furcata
Wiedemann 1819 must be examined to ensure that one does not represent the putative
new species. This is complicated by the fact that the putative new Nearctic species is
supported largely by DNA data. Until morphological characters can be found to support this
new taxon, there is no point in checking type specimens. The Neoascia and Xylota species
are straightforward but will be described by Chris Thompson in a planned publication.

New species to be described herein are in the genera Anasimyia Schiner 1864, Brachyopa
Meigen 1822, Hammerschmidtia Schummel 1834, Microdon Meigen 1803, Mixogaster
Macquart 1842, Neoascia, Orthonevra Macquart 1829, Pesilota Kieffer 1906 and
Trichopsomyia Williston 1888. Thompson (1991) studied the Nearctic taxa as part of a plan
to publish a Conspectus of Nearctic Syrphidae (hereafter, the “Conspectus”). Although
never published, this work has been privately distributed and has served as the backbone
of many taxonomic publications, including the Field Guide (Skevington et al. 2019).
Thompson and Vockeroth studied most types in preparing the Conspectus and it thus
provides a clear indication of taxonomic gaps for the family. Six of the species described
herein were first recorded in the Conspectus. Five of the 11 new species were discovered
during the preparation of the Field Guide.

Materials and methods

Adult specimens were pinned directly after collecting or stored in ethanol and later critical-
point dried in the case of Malaise trap samples. Clearing and removal of male genitalia
were necessary in order to view internal characters, useful for classification and species
delimitation. The genitalia capsule was removed by cutting between tergites 7 and 8 and
then cleared by heating in 88% lactic acid overnight. A full materials examined list is
provided after each species description. Specimens were borrowed from the following
institutions and individuals (collection codes based on Evenhuis (2007) are used in the
material examined sections):

4 Skevington J et al

ANSP - Academy of Natural Sciences of Drexel University, Philadelphia, Pennsylvania,
USA

CAS - California Academy of Sciences, San Francisco, California, USA
CBG - Centre for Biodiversity Genomics, Guelph, Ontario, Canada

CNC - Canadian National Collection of Insects, Arachnics, and Nematodes, Ottawa,
Ontario, Canada

DEBU - University of Guelph Insect Collection, Guelph, Ontario, Canada

MCZ - Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts
MEMU - Mississippi State University, Starkville, Mississippi, USA

NBMB - New Brunswick Museum, St. John's, New Brunswick

USNM - National Museum of Natural History, Washington D.C., USA

Photography

Habitus and genitalia photographs for all specimens were taken using a Leica M205-C
stereoscope (Leica Microsystems Inc., Concord, Ontario, Canada) using 0.6x (habitus) and
1.6x (genitalia) lenses. Raw images to be used in depth-of-field photomontages were
captured using Leica Application Suite (Leica Microsystems 2019) and final images were
created using Zerene Stacker (Littlefield 2018). Specimen measurements were taken using
the Leica measurement module in Leica Application Suite. Figures are presented
alphabetically.

DNA Sequencing

The right hind leg was removed from all recently collected (post 2000) specimens and sent
to the University of Guelph Biodiversity Institute of Ontario for sequencing of the 5' end of
the Cyctochrome c Oxidase | mitochondrial gene (COI) or Barcoding region, following
protocols published in Hajibabaei et al. (2005). DNA extraction and sequencing from older
specimens (most) was performed by Scott Kelso at the CNC using a modified version of
the same protocol, with custom primers developed for use in Diptera. These custom
primers (Table 1) sequence the Barcoding region in three smaller segments to be
assembled as a full barcode, allowing fragmented DNA from older specimens and
specimens stored in suboptimal conditions to be successfully sequenced. All sequence
data are stored on the BOLD website (www.boldsysiems.org) in the 'New Species of
Syrphidae from Field Guide’ (DS-NSSFG) dataset, available at dx.doi.org/10.5883/DS-
NSSEG. GenBank Numbers and unique identifiers (‘Sample ID') of specimens sequenced
are available in Table 2. With material sequenced at the CNC, raw sequence reads were
scored using Sequencher 5.4.6 (Gene Codes Corporation 2019) and aligned using
Mesquite (Maddison and Maddison 2010).

Table 1.

Primers used to sequence COI in this study.

Primer Name

New Syrphidae (Diptera) of North-eastern North America

Primer Design

Primer Sequence

Heb-F Folmer et al. 1994 GGT CAA CAAATC ATAAAG ATA TTG G
COI-Fx-A-R Young et al. (in prep) CGD GGR AAD GCY ATR TCD GG
COI-Fx-B-F Young et al. (in prep) GGD KCH CCN GAY ATR GC
COI-Fx-B-R Young et al. (in prep) GWA ATR AAR TTW ACD GCH CC
COI-Fx-C-F Young et al. (in prep) GGD ATW TCH TCH ATY YTA GG
COI-780R Gibson et al. 2011 CCAAAA AAT CAR AAT ARR TGY TG
Table 2.

Sequenced Specimens.

Species Sample ID Deposition GenBank Number Sequence Length
Anasimyia diffusa 10BBCDIP-0779 CBG JF866884 633
Anasimyia diffusa 10PROBE-13676 CNC JF877353 658
Anasimyia diffusa BIOUG04123-H06 CBG KM946129 592
Anasimyia diffusa BIOUG06854-C07 CBG KM936389 567
Anasimyia diffusa CNC_Diptera102200 CNC MNO015555 609
Anasimyia diffusa CNC_Diptera102201 CNC MNO15565 658
Anasimyia diffusa CNC_Diptera106474 CNC MN015574 375
Anasimyia diffusa CNC_Diptera44629 CNC MNO15577 583
Anasimyia diffusa CNC640314 CNC MNO15578 632
Anasimyia diffusa CNC640356 CNC MNO015572 633
Anasimyia diffusa CNC640369 CNC MNO015576 630
Anasimyia diffusa CNC640425 CNC MNO15557 631
Anasimyia diffusa CNC_Diptera4972 CNC MN015558 576
Anasimyia diffusa CNC_Diptera4974 CNC MNO15559 575
Brachyopa caesariata BlIOUG01347-D12 CBG KT104320 658
Brachyopa caesariata BlOUG04288-F04 CBG KR429817 658
Brachyopa caesariata BIOUG04346-B01 CBG KR433288 658
Brachyopa caesariata BlOUG04570-D11 CBG KR604654 670

Brachyopa caesariata BlOUG05447-E11 CBG KM950619 641

6 Skevington J et al

Species Sample ID Deposition GenBank Number Sequence Length
Brachyopa caesariata BIOUG05966-A03 CBG KM951714 617
Brachyopa caesariata BlOUG20323-G12 CBG ME837949 576
Brachyopa caesariata BlIOUG22325-H09 CBG KT607903 564
Brachyopa caesariata CNC_Diptera162914 CNC MNO015570 407
Brachyopa caesariata CNC_Diptera37551 CNC MNO15564 658
Hammerschmidtia sedmani BlOUG08107-B04 CBG KM958957 661
Hammerschmidtia sedmani BlOUG24033-B11 CBG KT 707288 506
Hammerschmidtia sedmani CNC_Diptera49268 CNC MNO15556 658
Hammerschmidtia sedmani CNC_Diptera49269 CNC MNO15560 658
Hammerschmidtia sedmani CNC_Diptera49270 CNC MNO15571 658
Hammerschmidtia sedmani + CNC298243 CNC MNO15575 646
Hammerschmidtia sedmani JKO0736 CNC MNO15573 658
Microdon scauros Jeff_Skevington_Specimen44177 ANSP MNO15566 667
Mixogaster fattigi Jeff_Skevington_Specimen45174 ANSP MNO15561 660
Neoascia guttata CNC_Dipteral69737 CNC MNO15562 667
Neoascia guttata CNC_Diptera170046 CNC MNO15567 434
Psilota klymkoi JK5333 CNC MNO0O15569 646
Trichopsomyia litoralis CNC_Diptera246379 CNC MNO15568 247
Terminology

Syrphidae-specific terminology follows Thompson (1999), while all other morphological
terminology follows Cumming and Wood (2017). A visual glossary of common terminology
used is available in Skevington et al. (2019).

Species Authorship

Authorship of each species is noted in the species description. As the species described
below were discovered and described by different people, not all of the authors of this
paper contributed equally to each species. In some cases, species discovered by now
deceased entomologists are attributed in part to them (depending on how many notes they
left on the taxon). In all cases, new data have been brought to bear on the species
concepts, including but not limited to genitalic morphology and DNA. Additional authors are
added to these species descriptions, based on their relative contributions.

New Syrphidae (Diptera) of North-eastern North America 7

Notes on Identification

All species included here can be identified by using Skevington et al. (2019). Keys are thus
not provided.

Taxon treatments
Anasimyia diffusa Locke, Skevington and Vockeroth, sp. n.

e ZooBank urn:lsid:zoobank.org:act:BF194AD1-F759-4EFO-BBB1-F8DD2EF53C7E

Nomenclature
Lejops (Anasimyia) diffusus Vockeroth manuscript name, in Thompson (1991)
Anasimyia undescribed species 1 Skevington et al. (2019), page 78

Materials

Holotype:

a. scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Ontario; locality: Oliver Bog, 3km, S Galt; decimalLatitude: 43.3168;
decimalLongitude: -80.283; eventDate: 1988-05-03/26; year: 1988; month: 5; day: 3;
verbatimEventDate: 3-26.v.1988; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera102201; recordedBy: D.Blades; identifiedBy: Michelle M.
Locke; dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

Paratypes:

a. scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Nunavut; locality: Akimiski Island; decimalLatitude: 53.105;
decimalLongitude: -80.957; eventDate: 2013-07; year: 2013; month: 7;
verbatimEventDate: viil.2013; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC640314; recordedBy: D. Beresford; identifiedBy: Michelle M.
Locke; dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

b. scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Nunavut; locality: Akimiski Island; decimalLatitude: 53.105;

Skevington J et al

decimalLongitude: -80.957; eventDate: 2013-07; year: 2013; month: 7;
verbatimEventDate: vii.2013; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC640356; recordedBy: D. Beresford; identifiedBy: Michelle M.
Locke; dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Ontario; locality: District: 2E-1 Albany River; decimalLatitude: 51.6492;
decimalLongitude: -81.848522; samplingProtocol: Malaise trap; eventDate:
2012-06-17; year: 2012; month: 6; day: 17; verbatimEventDate: 17.vi.2012; habitat:
Forest, Closed Treed cover, Eco District 2E-1 Albany River; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: CNC640369; recordedBy: Alex Howard &
Stan Vasilauskas; otherCatalogNumbers: 1101721B-2012-088-00008; identifiedBy:
Michelle M. Locke; dateldentified: 2019; language: en; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Ontario; locality: District: 1E-3 Winisk River; decimalLatitude: 54.4712;
decimalLongitude: -88.5649; samplingProtocol: Malaise trap; eventDate: 2011-06-21;
year: 2011; month: 6; day: 21; verbatimEventDate: 21.vi.2011; habitat: Forest,
Bryophytes, Herbaceous plant cover, Eco District 1E-3 Winisk River; individualCount:
1; sex: male; lifeStage: adult; catalogNumber: CNC640425; recordedBy: lan Fife &
Stan Phippen; otherCatalogNumbers: 1094706-2011-089-00002; identifiedBy: Michelle
M. Locke; dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Nunavut; locality: Akimiski Island; decimalLatitude: 53.105;
decimalLongitude: -80.957; samplingProtocol: Malaise trap; eventDate: 2017-07-17;
year: 2017; month: 7; day: 17; verbatimEventDate: 17.vii.2017; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC641362; recordedBy: David Beresford;
identifiedBy: Michelle M. Locke; dateldentified: 2019; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Nunavut; locality: Akimiski Island; decimalLatitude: 53.105;
decimalLongitude: -80.957; samplingProtocol: Malaise trap; eventDate: 2014-07-21;
year: 2014; month: 7; day: 21; verbatimEventDate: 21.vii.2014; individualCount: 1; sex:

New Syrphidae (Diptera) of North-eastern North America ss)

male; lifeStage: adult; catalogNumber: CNC641363; recordedBy: David Beresford;
otherCatalogNumbers: 30/27; identifiedBy: Michelle M. Locke; dateldentified: 2019;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Nunavut; locality: Akimiski Island; decimalLatitude: 53.105;
decimalLongitude: -80.957; samplingProtocol: Malaise trap; eventDate: 2014-07-21;
year: 2014; month: 7; day: 21; verbatimEventDate: 21.vii.2014; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC641364; recordedBy: David Beresford;
otherCatalogNumbers: 45/27; identifiedBy: Michelle M. Locke; dateldentified: 2019;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: New Brunswick; locality: Kouchibouguac National Park;
decimalLatitude: 46.85; decimalLongitude: -64.97; eventDate: 1978-06-15; year: 1978;
month: 6; day: 15; verbatimEventDate: 15.vi.1978; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera102200; recordedBy: S.J.Miller;
identifiedBy: Michelle M. Locke; dateldentified: 2019; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: British Columbia; locality: 6 km E[ast of] Salmon Arm; decimalLatitude:
50.7; decimalLongitude: -119.3; eventDate: 1992-06-30; year: 1992; month: 4; day: 30;
verbatimEventDate: 30.iv.1992; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera245059; recordedBy: A. Borkent; identifiedBy: Michelle
M. Locke; dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Manitoba; locality: O'day; decimalLatitude: 57.582; decimalLongitude:
-94.069; samplingProtocol: Malaise trap; eventDate: 2011-07-07/19; year: 2011;
month: 7; day: 7; verbatimEventDate: 7-19.viil.2011; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera284501; recordedBy: CANPOLIN;
otherCatalogNumbers: UG: SYRPH: 10763; identifiedBy: Michelle M. Locke;
dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:

10

Skevington J et al

Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Manitoba; locality: Herchmer; decimalLatitude: 57.378;
decimalLongitude: -94.195; samplingProtocol: Malaise trap; eventDate: 2011-07-07/19;
year: 2011; month: 7; day: 7; verbatimEventDate: 7-19.vii.2011; individualCount: 1;
sex: male; lifeStage: adult; catalogNumber: CNC_Diptera284505; recordedBy:
CANPOLIN; otherCatalogNumbers: UG: SYRPH: 01075; identifiedBy: Michelle M.
Locke; dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Alberta; locality: Waterton Lakes National Park; decimalLatitude:
49.083314; decimalLongitude: -113.916733; eventDate: 1980-07-07/12; year: 1980;
month: 7; day: 7; verbatimEventDate: 7-12.vii.1980; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera44602; recordedBy: H.J. Teskey;
identifiedBy: Michelle M. Locke; dateldentified: 2019; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: British Columbia; locality: 6 km East of Salmon Arm; decimalLatitude:
50.7; decimalLongitude: -119.2; eventDate: 1990-05-16; year: 1990; month: 5; day: 16;
verbatimEventDate: 16.v.1990; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera44629; recordedBy: A. Borkent; identifiedBy: Michelle M.
Locke; dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: British Columbia; locality: 6 km East of Salmon Arm; decimalLatitude:
50.7; decimalLongitude: -119.2; eventDate: 1990-05-16; year: 1990; month: 5; day: 16;
verbatimEventDate: 16.v.1990; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera44630; recordedBy: A. Borkent; identifiedBy: Michelle M.
Locke; dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Ontario; locality: Carleton Co. 8 km S Richmond fen; decimalLatitude:
45.069536; decimalLongitude: -75.854558; eventDate: 1983-08-02; year: 1983; month:

New Syrphidae (Diptera) of North-eastern North America 11

8; day: 2; verbatimEventDate: 2.viii.1983; individualCount: 1; sex: female; lifeStage:
adult; catalogNumber: CNC_Diptera4972; recordedBy: Cooper, Wood; identifiedBy:
Michelle M. Locke; dateldentified: 2019; language: en; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: New Brunswick; locality: Kouchibouguac National Park;
decimalLatitude: 46.85; decimalLongitude: -64.97; eventDate: 1978-06-14; year: 1978;
month: 6; day: 14; verbatimEventDate: 14.vi.1978; individualCount: 1; sex: female;
lifeStage: adult; catalogNumber: CNC_Diptera4974; recordedBy: S.J. Miller;
identifiedBy: Michelle M. Locke; dateldentified: 2019; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Manitoba; locality: Aweme; decimalLatitude: 49.7; decimalLongitude:
-99.6; eventDate: 1907-08-25; year: 1907; month: 8; day: 25; verbatimEventDate:
25.vill.L907; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC_Diptera91228; recordedBy: J. Fletcher; identifiedBy: Michelle M. Locke;
dateldentified: 2019; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Ontario; locality: Oliver Bog, 3km, S Galt; decimalLatitude: 43.3168;
decimalLongitude: -80.283; samplingProtocol: pan trap; eventDate: 1988-05-03/26;
year: 1988; month: 5; day: 3; verbatimEventDate: 3-26.v.1988; habitat: pt2 expanse
hollow; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen17433; recordedBy: D. Blades; otherCatalogNumbers:
accession#026W; identifiedBy: Michelle M. Locke; dateldentified: 2019; language: en;
institutionCode: DEBU; basisOfRecord: PreservedSpecimen

scientificName: Anasimyia diffusa Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Smooth-legged Swamp Fly; genus: Anasimyia; specificEpithet: diffusa;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: Canada;
stateProvince: Ontario; locality: Oliver Bog, 3km, S Galt; decimalLatitude: 43.3168;
decimalLongitude: -80.283; eventDate: 1988-05-03/26; year: 1988; month: 5; day: 3;
verbatimEventDate: 3-26.v.1988; habitat: pt2 expanse hollow; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen17434; recordedBy:
D. Blades; otherCatalogNumbers: accession#026W; identifiedBy: Michelle M. Locke;
dateldentified: 2019; language: en; institutionCode: DEBU; basisOfRecord:
PreservedSpecimen

12

Skevington J et al

Description
Size: Body length 7.6 to 10.7 mm; wing length 5.6 to 7.4 mm

Male: Head: Face yellow, covered in yellow pollen and fine yellow pile, sides of face
black, shining and bare, small medial tubercle, lower face projecting forwards along the
oral margin; gena black, covered in yellow pollen and fine yellow pile; frons black
dorsally, yellow ventrally, sometimes narrowly so just along ventral edge and across
dorsum of antennal socket, dense yellow pollen often obscures ground colour, narrow
shining spot dorsal to the antennal socket, yellow pilose, sometimes with a few black
pili; vertex black, mostly black pilose with some yellow pile; antenna wholly yellow to
orange, scape and pedicel with short, black pile, pedicel slightly longer than scape,
postpedicel round; eye bare.

Thorax: Scutum dull black with two medial, narrow, yellow pollinose stripes, lateral
edges yellow pollinose from postpronotum to postalar callus, yellow _ pilose;
postpronotum black, covered in yellow pollen and yellow pile; postalar callus black,
covered in yellow pollen and yellow pile; scutellum yellow with black lateral edges and
narrowly black along anterior edge, yellow pilose with some black pili admixed; pleuron
black, covered in yellow pollen, anterior anepisternum, katepimeron and meron bare,
posterior anepisternum, katepisternum and anepimeron yellow pilose; metasternum
black, yellow pollinose, yellow pilose; pro- and mesolegs with coxae and trochanters
black, yellow pilose, femora black on basal %, yellow on apical %, black extends almost
to apex anteroventrally, sometimes more or less extensively black, anteroventral basal
patch of black setulae, yellow pilose; protibia yellow, usually brown to black anteriorly
on apical %4-'/3, covered in short, yellow pile, protarsus brown anteriorly, yellow
posteriorly, similar in colour to apex of protibia, covered in short yellow pile, few black
pili anteriorly; mesotibia yellow, sometimes brown, yellow pilose; mesotarsus yellow
basally, brown to black apically, tarsomeres 1 and 2 yellow, 3 yellow or brown, 4 and 5
brown, all yellow pilose with black setulae posteriorly; metaleg with coxa black, yellow
pollinose, yellow pilose; metatrochanter without tubercle, black, shining, long yellow
pile anteriorly, some short yellow pile posteriorly; metafemur enlarged, yellow,
anteromedial spot ranging from obscure brown to black spot ~’% the length of the
metafemur to large black spot on medial 2 of femur, posterior side 7/3-% black medially,
dorsally extending to apex, apical end of metafemur narrowly black, metafemur yellow
pilose, small patch of black setulae anterobasally, black setulae along ventral side;
metatibia brown to black basally and apically, yellow on medial %4-'/, with ventral
carinate ridge, yellow pilose, apical end projected forwards slightly beyond end of tibia
not formed into noticeable spur; metatarsus brown to black, black pilose anteriorly,
yellow pilose posteriorly; wing entirely densely microtrichose; halter yellow; calypter
yellow.

Abdomen: Black, yellow pilose with few black pili admixed; tergite 1 pollinose
anterolaterally; tergite 2 yellow laterally, widely on anterior 3%4-%s, usually narrowly on
posterior %s-%, yellow extending medially on to tergite and tapering to a point about %4-
'/s from lateral edge, yellow cuticle covered in yellow pollen, sometimes extending

New Syrphidae (Diptera) of North-eastern North America 13

slightly on to black cuticle medially where it tapers to a point; tergite 3 with anterolateral
yellow spots % length of tergite, often narrowly yellow laterally on posterior half, pair of
yellow to grey curved pollinose markings medially; tergite 4 similar to tergite 3 but
lateral edges only narrowly yellow, no anterolateral yellow spots, narrowly yellow
posteriorly (tergite 3 sometimes very narrowly yellow posteriorly as well); tergites 2-4
shining on posterior ’/s-12, typically more widely shining on tergites 3 and 4; sternite 1
black on anterior 34, yellow on posterior 14, black area convex; sternite 2 yellow, often
with brown to black spot posteromedially, spot often obscure, sternite 3 often brown to
black with posterior edge narrowly yellow, sometimes yellow on anterior 42; sternite 4
brown to black with posterior edge narrowly yellow; all sternites yellow to grey pollinose
with yellow pile.

Genitalia: Epandrium longer than wide, narrows slightly dorsally, with apicodorsal ridge
adjacent to cerci; cercus with laterally compressed, subquadrate, sclerotised outer
portion covered in long pile and membranous inner portion; surstylus long and wide,
bent, with small patch of setae on ventral inner surface at bend, apical end bilobed and
curved inwards, dorsal and ventral lobes covered in setae; basal hypandrium convexly
curved ventrally, with two low, parallel ridges ending with several posterior-facing,
curved setae at apicoventral end (one pair of setae larger than other setae); apical
hypandrium split into two complex, multi-lobed arms; phallapodeme long, rod-like
structure, laterally compressed and curved dorsally at base; phallus is difficult to
distinguish, hidden between the multi-lobed hypandrial arms, small, short, with two
pairs of lobes, basal lobes broad and directed laterally, distal lobes narrow and directed
apically, with separate dorsal conical structure, pointed at apex (possibly secondarily
derived from surrounding membrane).

Female: Differs from male in the following ways: frons black with narrow yellow ventral
edge including dorsum of antennal socket, black pilose; scutellum often entirely yellow
pilose, sometimes with few black pili admixed; pro- and mesofemora black on basal ¥/3-
4%, yellow on apical 42-7, black extends almost to apex ventrally, sometimes less
extensively black basally with black not extending entirely around femur; metafemur
with brown anterior spot smaller than in male, sometimes completely absent; metatibia
similar to male but often more yellow with brown areas reduced especially basally;
abdomen with short subappressed yellow and black pile; tergite 1 entirely pollinose,
with short, yellow subappressed pile; tergite 2 yellow anterolaterally, sometimes with
yellow extending narrowly along lateral margin, with pair of curved pollinose markings
beginning anterolaterally and curving inwards medially, pile subappressed with some
long pile on anterolateral edge, yellow pilose on anterior % or more, black pilose on
posterior % or less, posterior edge shining; tergites 3 and 4 similar to tergite 2,
sometimes with posterior edge narrowly yellow and with small medial pollinose marking
posteromedially, often more extensively black pilose on posterior % or less, pile
uniformly short, no long pile anterolaterally, tergites 3 and 4 not as long as tergite 2;
tergite 5 smaller than other tergites, black anteriorly, yellow posteriorly, tergite almost
entirely pollinose except along anterior edge and narrow medial stripe, yellow pilose
with few black pili posteriorly; sternites pollinose and yellow pilose; sternite one black;

14

Skevington J et al

sternite 2 mostly yellow with black medial triangular marking; sternites 3 and 4 mostly
black with narrow yellow lateral and sometimes posterior edges.

See Fig. 1 & Fig. 2.

a

Figure 1.

Anasimyia diffusa sp. nov.

a: Dorsal habitus, CNC_Diptera44630 EES]

b: Lateral habitus, CNC_Diptera44630 EES

c: Face, oblique, Holotype, CNC_Diptera102201 EES
d: Metaleg, lateral, Holotype, CNC_Diptera102201 ERS]
e: Metaleg, ventral, Holotype, CNC_Diptera102201 ER

New Syrphidae (Diptera) of North-eastern North America 15

Figure 2.

Anasimyia diffusa sp. nov., male genitalia, scale bars all 200 um

a: Lateral terminalia, Jeff_Skevington_Specimen17434 ETS]

b: Ventral terminalia, Jeff_Skevington_Specimen17434 ETS]

c: Lateral hypandrium, Jeff_Skevington_Specimen17434 EQS]

d: Ventral of hypandrium, Jeff_Skevington_Specimen17434 EE]

Diagnosis

Male metatrochanter without tubercle, with fine, yellow pile anteriorly (Fig. 1e);
metatibia without spur (Fig. 1d) and tergite 4 with large pair of curved pollinose
markings.

Etymology

The word diffusa is Latin, meaning spread out, extended, dispersed. This is the name
given to this species by J.R. Vockeroth, presumably with reference to the wide
distribution.

Distribution

This species is known from 20 specimens from New Brunswick to British Columbia.

16 Skevington J et al

Ecology

Anasimyia diffusa is often collected in the same location as the much more common A.
anausis (Walker 1849). They appear to both have a broad distribution and a long flight
season, with A. diffusa collected from mid-April to late August. Some specimens have
been collected in bogs but it is unclear if it is a bog specialist or not.

Taxon discussion

Differences in the metatrochanter are the easiest way to distinguish A. diffusa from A.
anausis. The metatrochanter of A. anausis has short black setuale anteroventrally. In
some specimens these setulae appear to be broken off but their insertions into the
cuticle give the metatrochanter a rough texture. Anasimyia bilinearis (Williston 1887)
can be differentiated from A. diffusa by its mostly dark abdomen with reduced pollinose
markings. There is no tubercle on the metatrochanter of A. diffusa as there is in
Anasimyia Group 2 males (See Skevington et al. 2019). Females are similar to other
Group 1 females (See Skevington et al. 2019) and cannot be distinguished from similar
species. Five females of A. diffusa were identified using the barcoding region of COI.
These specimens in the paratype series clustered in a neighbour-joining tree with four
males that were morphologically identified to A. diffusa. DNA sequences of this species
are available in the public BOLD dataset "New Species of Syrphidae from Field Guide -
DS-NSSFG" and in GenBank (Table 2). They are genetically most similar to A. anausis
but consistently separable with DNA barcodes.

Common Name

The common name given to the species by Skevington et al. (2019) is Smooth-legged
Swamp Fly.

Anasimyia matutina Locke, Skevington and Vockeroth, sp. n.

° ZooBank urn:|sid:zoobank.org:act:2FCCCB1B-FBA1-4E33-BAE6-F5D722A5ECF8

Nomenclature
Lejops (Anasimyia) matutinus Vockeroth manuscript name, in Thompson (1991)
Anasimyia undescribed species 2 Skevington et al. (2019), page 80

Materials

Holotype:

a. scientificName: Anasimyia matutina Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Small-spotted Swamp Fly; genus: Anasimyia; specificEpithet: matutina;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: U.S.A.;

New Syrphidae (Diptera) of North-eastern North America

stateProvince: Virginia; locality: Great Falls; decimalLatitude: 39; decimalLongitude:
-77.283333; samplingProtocol: hand collected; eventDate: 06/16/1952; year: 1952:
month: 6; day: 16; verbatimEventDate: 16.vi.1952; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera91232; recordedBy: N. Banks;
identifiedBy: J.R. Vockeroth; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

17

Paratypes:

a. scientificName: Anasimyia matutina Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Small-spotted Swamp Fly; genus: Anasimyia; specificEpithet: matutina;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: U.S.A.;
stateProvince: Virginia; locality: Petersburg, Chesterford County; decimalLatitude:
37.227858; decimalLongitude: -77.40195; samplingProtocol: hand collected;
eventDate: 06/01/1917; year: 1917; month: 6; day: 1; verbatimEventDate: 1.vi.1917;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera91233;
identifiedBy: J.R. Vockeroth; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

b. scientificName: Anasimyia matutina Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Small-spotted Swamp Fly; genus: Anasimyia; specificEpithet: matutina;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: U.S.A.;
stateProvince: Washington, D.C.; locality: Washington; decimalLatitude: 38.9;
decimalLongitude: -77.033333; samplingProtocol: hand collected; eventDate:
05/17/1912: year: 1912; month: 5; day: 17; verbatimEventDate: 17.v.1912;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera91234;
recordedBy: Fred K. Knab; identifiedBy: J.R. Vockeroth; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

C: scientificName: Anasimyia matutina Locke, Skevington and Vockeroth;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Small-spotted Swamp Fly; genus: Anasimyia; specificEpithet: matutina;
scientificNameAuthorship: Locke, Skevington and Vockeroth; country: U.S.A.;
stateProvince: Virginia; locality: Chain Bridge; decimalLatitude: 38.929911:
decimalLongitude: -77.114411; samplingProtocol: hand collected; eventDate: 06-14;
month: 6; day: 14; verbatimEventDate: 14.vi; individualCount: 1; sex: male; lifeStage:
adult; catalogNumber: Jeff_Skevington_Specimen17432; recordedBy: N. Banks;
identifiedBy: J.R. Vockeroth; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

Description

Size: Body length 9.8 to 10.2 mm; wing length 6.1 to 6.4 mm

Male: Head: Face yellow, covered in yellow pollen and fine yellow pile, sides of face
black, shining and bare, small medial tubercle, lower face projecting forwards along the
oral margin; gena black, covered in yellow pollen and fine yellow pile; frons black
dorsally, yellow ventrally, dense yellow pollen often obscures ground colour, narrow

18

Skevington J et al

shining spot dorsal to the antennal socket, yellow pilose; vertex black, mostly black
pilose with some yellow pile; antenna wholly yellow to orange, scape and pedicel with
short, black pile, pedicel slightly longer than scape, postpedicel round; eye bare.

Thorax: Scutum dull black with two medial, yellow pollinose stripes that are more than
1/ as wide as adjacent black area, lateral edges yellow pollinose from postpronotum to
postalar callus, yellow pilose; postpronotum black, covered in yellow pollen and yellow
pile; postalar callus black, covered in yellow pollen and yellow pile; scutellum black on
basal %, yellow on apical %, yellow pilose; pleuron black, covered in yellow pollen,
anterior anepisternum, katepimeron and meron bare, posterior anepisternum,
katepisternum and anepimeron yellow pilose; metasternum black, yellow pollinose,
yellow pilose; procoxa black, yellow pollinose, yellow pilose; protrochanter black, yellow
pollinose, yellow pilose with a few black setulae apically; profemur black on basal 12-74,
yellow on apical -%%, yellow pollinose, yellow pilose, anteroventral basal patch of
black setulae with cuticle shining below; protibia yellow, yellow pilose; protarsus yellow,
yellow pilose, apical two tarsomeres sometimes darkened brown; mesocoxa black,
yellow pollinose, yellow pilose, few black setulae apically; mesotrochanter black, yellow
pollinose, yellow pilose, few black setulae apically; mesofemur black on basal %,
yellow on apical %, yellow pollinose, yellow pilose, anteroventral basal patch of black
setulae; mesotibia yellow, yellow pilose with few black setulae ventrally on apical end;
mesotarsus yellow, yellow pilose with black setulae ventrally, apical one or two
tarsomeres sometimes slightly darkened brown; metacoxa black, yellow pollinose,
yellow pilose anteriorly; metatrochanter black, yellow pilose with small ventral tubercle;
metafemur enlarged, black basal 7, yellow apical ’/3 with apex narrowly black, yellow
pilose, black setulae ventrally on apical 7; metatibia light brown, sometimes yellow, on
basal '/s, yellow on medial ’/3, dark brown (darker than base) on apical ’/3 (Sometimes
>'/3), yellow pilose, with ventral carinate ridge ending in long apical spur, spur %4-’/s as
long as basotarsomere; metatarsus brown, black pilose anteriorly, yellow pilose
posteriorly; wing entirely densely microtrichose; halter yellow; calypter yellow.

Abdomen: Tergite 1 black, small yellow spots on lateral edges, covered in yellow to
grey pollen, yellow pilose; tergite 2 elongate, longer than wide and longer than tergite
3, slightly narrowing posteriorly, yellow laterally, black medially forming a ‘v’ shape that
is widest anteriorly, narrowest medially, posterior % brown to black, pair of faint yellow
pollinose markings on yellow cuticle that extend from anterolateral edge and curve
inwards medially, yellow pilose, posterior edge shining; tergite 3 brown to reddish to
black, sometimes more yellow to orange, usually not uniform in colour and without a
distinct pattern, anterolateral corners yellow with yellow pollen, yellow pilose, posterior
edge shining; tergite 4 similar to tergite 3 but usually with anterior medial pair of small
pollinose spots, faint pollinose spot between dull anterior portion and shining posterior
margin; sternite 1 black, yellow to grey pollinose, yellow pilose; sternite 2 yellow
anteriorly, black posteriorly, black extends to or almost to anterior edge in a narrow,
medial point; sternite 3 brown to black anteriorly, yellow to brown posteriorly, amount
varies from % black to almost entirely black; sternite 4 yellow to brown; sternites 2-4

New Syrphidae (Diptera) of North-eastern North America 19

covered in yellow to grey pollen, pollen is more dense, often forming bands, along the
posterior edge, yellow pilose.

Genitalia: Epandrium subquadrate, longer than wide, narrows slightly dorsally; cercus
with laterally compressed, subquadrate sclerotised outer portion covered in long pile
and membranous inner portion; surstylus long, bent at a 90° angle, with setae on
ventral inner surface, apical end triangular, curved inwards, covered in setae on dorsal
and ventral edges; basal hypandrium with central, basoventral pointed process,
directed apically, narrows medially, deeply concave ventrally, with several
posteroventral-facing curved setae at apicoventral end (one pair of setae longer than
other setae); apical hypandrium split into two complex, multi-lobed arms; phallapodeme
long, rod-like, laterally compressed and curved dorsally at base; phallus situated
between the multi-lobed hypandrial arms, small and difficult to distinguish, with
separate small, dorsal conical structure, pointed at apex (possibly secondarily derived
from surrounding membrane).

Female: Unknown.

See Fig. 3 & Fig. 4.

a

Yds 7 =
A edd a
WT) Ne inn

Figure 3.

Anasimyia matutina sp. nov.

a: Dorsal habitus, Holotype, CNC_Diptera91232 ER

b: Lateral habitus, Jeff_Skevington_Specimen17432 ETS]
c: Head, oblique, Holotype, CNC_Diptera91232 [ETS]

d: Hind leg, Jeff_Skevington_Specimen17432 EE]

Skevington J et al

Figure 4.
Anasimyia matutina sp. nov., male genitalia

a: Lateral of male terminalia, Jeff_Skevington_Specimen17432; scale bar 500 um ES]
b: Ventral of male terminalia, Jeff_Skevington_Specimen17432; scale bar 200 um EES]
c: Sperm pump and ejaculatory apodeme, Jeff_Skevington_Specimen17432; scale bar 100

um EE
d: Ventral of hypandrium, Jeff_Skevington_Specimen17432; scale bar 200um EES]
Diagnosis

Male metatrochanter with modest tubercle and long, acute apical metatibial spur (Fig.
3d).

Etymology

The word matutina is Latin, meaning of the morning, early. This is the name given to
this species by J.R. Vockeroth.

Distribution

Only four specimens are known from Virginia (Chain Bridge, Great Falls and
Petersburg) and Washington DC.

New Syrphidae (Diptera) of North-eastern North America 21

Ecology

Rarely found, therefore nothing is known about its habitat association. It is known to fly
from mid-May to mid-June.

Taxon discussion

Size and shape of spur on metatrochanter and of metatibial spur separate this from
other Anasimyia Group 2 males (see Skevington et al. 2019). Females are unknown,
but are likely to be identifiable as Anasimyia Group 2 females (Skevington et al. 2019).

Common Name

The common name given to the species by Skevington et al. (2019) is Small-spotted
Swamp Fly.

Brachyopa caesariata Moran and Skevington, sp. n.

° ZooBank urn:lsid:zoobank.org:act:A39EOE80-49A9-41F9-877A-D9B3AF5A2347

Nomenclature
Brachyopa 78-2 Thompson manuscript name, in Thompson (1991)

Brachyopa undescribed species 78-2 Skevington et al. (2019), page 210

Materials

Holotype:

a. scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Algonquin Provincial
Park, Opeongo Road; decimalLatitude: 45.625911; decimalLongitude: -78.350753;
eventDate: 06/03/2018; year: 2018; month: 6; day: 3; verbatimEventDate: 3.vi.2018;
habitat: on flowers of Prunus virginiana; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC1042805; recordedBy: J.H.Skevington, M.M. Locke;
otherCatalogNumbers: JSM11211; identifiedBy: K. Moran; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

Paratypes:

a. scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; siateProvince: Ontario; locality: Algonquin Provincial
Park, Opeongo Road; decimalLatitude: 45.625911; decimalLongitude: -78.350753;
eventDate: 06/03/2018; year: 2018; month: 6; day: 3; verbatimEventDate: 3.vi.2018;

22

Skevington J et al

habitat: on flowers of Prunus virginiana; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC1042806; recordedBy: J.H.Skevington, M.M. Locke;
otherCatalogNumbers: JSM11212; identifiedBy: K. Moran; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Washington; locality: ValleyFord;
decimalLatitude: 47.535775; decimalLongitude: -117.238994; eventDate: 05/17/1924;
year: 1924; month: 5; day: 17; verbatimEventDate: 17.v.1924; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC1185344; recordedBy: A.L. Melander;
identifiedBy: F.C. Thompson; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: British Columbia; locality: Kamloops;
decimalLatitude: 50.674234; decimalLongitude: -120.299615; eventDate: 05/01/1932;
year: 1932; month: 5; day: 1; verbatimEventDate: 1.v.1932; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC1185345; recordedBy: O. Bryant;
identifiedBy: F.C. Thompson; language: en; institutionCode: CAS; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Washington; locality: Walla Walla;
decimalLatitude: 46.069657; decimalLongitude: -118.356384; eventDate: 04/12/1924;
year: 1924; month: 4; day: 12; verbatimEventDate: 12.iv.1924; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC1185346; recordedBy: V. Argo;
identifiedBy: F.C. Thompson; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Washington; locality: Asotin;
decimalLatitude: 46.341789; decimalLongitude: -117.055822; eventDate: 04/06/1924;
year: 1924; month: 4; day: 6; verbatimEventDate: 6.iv.1924; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC1185347; recordedBy: V. Argo;
otherCatalogNumbers: USNMENT01518169; identifiedBy: F.C. Thompson; language:
en; institutionCode: USNM; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Utah; locality: Cache Co., Logan Canyon,
Twin Creek; decimalLatitude: 41.867666; decimalLongitude: -111.594781; eventDate:

New Syrphidae (Diptera) of North-eastern North America 23

1979-06-12/21; startDayOfYear: 163; endDayOfYear: 172; year: 1979; month: 6; day:
12; verbatimEventDate: 12-21.vi.1979; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC1185348; identifiedBy: F.C. Thompson; language: en;
institutionCode: USNM; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Maynooth, Hastings
County; decimalLatitude: 45.230639; decimalLongitude: -77.938375; eventDate:
05/10/1987; year: 1987; month: 5; day: 10; verbatimEventDate: 10.v.1987;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera101644;
recordedBy: Dbell,M.Wood; identifiedBy: K. Moran; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Maynooth, Hastings
County; decimalLatitude: 45.230639; decimalLongitude: -77.938375; eventDate:
05/10/1987; year: 1987; month: 5; day: 10; verbatimEventDate: 10.v.1987;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera106863;
recordedBy: D. Bell, M. Wood; identifiedBy: K. Moran; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Alaska; locality: 11 Miles South Anderson
Junction, Mile 270 Rte 3; decimalLatitude: 64.165328; decimalLongitude: -149.287278;
samplingProtocol: Malaise trap; eventDate: 1984-06/08-23/11; startDayOfYear: 174;
endDayOfYear: 224; year: 1984; month: 6; day: 23; verbatimEventDate:
23.vi.-11.vili.1984; habitat: Alnus Populus-Picea; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Dipterai07450; recordedBy: S. & J. Peck;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Alaska; locality: 11 Miles South Anderson
Junction, Mile 270 Rte 3; decimalLatitude: 64.165328; decimalLongitude: -149.287278;
samplingProtocol: Malaise trap; eventDate: 1984-06/08-23/11; startDayOfYear: 174;
endDayOfYear: 224; year: 1984; month: 6; day: 23; verbatimEventDate:
23.vi.-11.vill.1984; habitat: Alnus Populus-Picea; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Dipterai07451; recordedBy: S. & J. Peck;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;

24

Skevington J et al

family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Alaska; locality: 11 Miles South Anderson
Junction, Mile 270 Rte 3; decimalLatitude: 64.165328; decimalLongitude: -149.287278;
samplingProtocol: Malaise trap; eventDate: 1984-06/08-23/11; startDayOfYear: 174;
endDayOfYear: 224; year: 1984; month: 6; day: 23; verbatimEventDate:
23.Vi.-11.vili.1984; habitat: Alnus Populus-Picea; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera107454; recordedBy: S. & J. Peck;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Alaska; locality: 11 Miles South Anderson
Junction, Mile 270 Rte 3; decimalLatitude: 64.165328; decimalLongitude: -149.287278;
samplingProtocol: Malaise trap; eventDate: 1984-06/08-23/11; startDayOfYear: 174;
endDayOfYear: 224; year: 1984; month: 6; day: 23; verbatimEventDate:
23.Vi.-11.vill.1984; habitat: Alnus Populus-Picea; individualCount: 1; sex: female;
lifeStage: adult; catalogNumber: CNC_Diptera107456; recordedBy: S. & J. Peck;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Ottawa, Marlborough
Forest Rideau Trail; decimalLatitude: 45.05; decimalLongitude: -75.816667; eventDate:
05/25/2011; year: 2011; month: 5; day: 25; verbatimEventDate: 25.v.2011; habitat:
near Rogers Pond; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC_Diptera243273; recordedBy: B.J. Sinclair; identifiedBy: K. Moran; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Quebec; locality: Laniel; decimalLatitude:
47.033333; decimalLongitude: -79.266667; eventDate: 06/10/1931; year: 1931; month:
6; day: 10; verbatimEventDate: 10.vi.1931; individualCount: 1; sex: male; lifeStage:
adult; catalogNumber: CNC_Diptera37403; recordedBy: M.B. Dunn; identifiedBy: K.
Moran; language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Maynooth;
decimalLatitude: 45.233333; decimalLongitude: -77.95; eventDate: 06/18/1953; year:
1953; month: 6; day: 18; verbatimEventDate: 18.vi.1953; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: CNC_Diptera37420; recordedBy: J.F.

New Syrphidae (Diptera) of North-eastern North America 25

McAlpine; identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Quebec; locality: Duncan Lake, near
Rupert; decimalLatitude: 45.681389; decimalLongitude: -76.050278; eventDate:
06/14/1969; year: 1969; month: 6; day: 14; verbatimEventDate: 14.vi.1969;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera37461;
recordedBy: J.F. McAlpine; identifiedBy: K. Moran; language: en; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: British Columbia; locality: Liard Hot
Springs, Mi. 496 Alaska Hwy; minimumElevationInMeters: 457; decimalLatitude:
59.425992; decimalLongitude: -126.0964; eventDate: 07/09/1959; year: 1959; month:
7; day: 9; verbatimEventDate: 9.vii.1959; individualCount: 1; sex: female; lifeStage:
adult; catalogNumber: CNC_Diptera37465; recordedBy: R.E. Leech; identifiedBy: K.
Moran; language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: British Columbia; locality: Liard Hot
Springs, Mi. 496 Alaska Hwy; minimumElevationInMeters: 457; decimalLatitude:
59.425992; decimalLongitude: -126.0964; eventDate: 07/09/1959; year: 1959; month:
7; day: 9; verbatimEventDate: 9.vii.1959; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera37547; recordedBy: R.E. Leech; identifiedBy: K. Moran;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Alberta; locality: Edmonton;
decimalLatitude: 53.540942; decimalLongitude: -113.4937; eventDate: 06/15/1948;
year: 1948; month: 6; day: 15; verbatimEventDate: 15.vi.1948; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC_Diptera37548; recordedBy: J.R.
Vockeroth; identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Mer Bleue Ottawa;
decimalLatitude: 45.4; decimalLongitude: -75.5; eventDate: 05/31/1923; year: 1923;
month: 5; day: 31; verbatimEventDate: 31.v.1923; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera37549; recordedBy: C.H. Curran;

26

Skevington J et al

identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Ottawa; decimalLatitude:
45.423; decimalLongitude: -75.698; eventDate: 06/02/1927; year: 1927; month: 6; day:
2; verbatimEventDate: 2.vi.1927; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera37550; recordedBy: C.H. Curran; identifiedBy: K. Moran;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Ottawa nr. Uplands
Airport; decimalLatitude: 45.333333; decimalLongitude: -75.583333; eventDate:
05/29/1990; year: 1990; month: 5; day: 29; verbatimEventDate: 29.v.1990;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera37551;
recordedBy: J.M. Cumming; identifiedBy: K. Moran; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Manitoba; locality: Aweme;
decimalLatitude: 49.708531; decimalLongitude: -99.602758; eventDate: 06/09/1925;
year: 1925; month: 6; day: 9; verbatimEventDate: 9.vi.1925; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC_Diptera37555; recordedBy: R.D. Bird;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Nova Scotia; locality: Kentville;
decimalLatitude: 45.066667; decimalLongitude: -64.483333; eventDate: 06/09/1915;
year: 1915; month: 6; day: 9; verbatimEventDate: 9.vi.1915; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC_Diptera37556; identifiedBy: K. Moran;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: British Columbia; locality: Hixon;
decimalLatitude: 53.420261; decimalLongitude: -122.585961; eventDate: 06/07/1966;
year: 1966; month: 6; day: 7; verbatimEventDate: 7.vi.1966; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC_Diptera37559; recordedBy: E.D.A. Dyer;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

aa.

ab.

ac.

ad.

New Syrphidae (Diptera) of North-eastern North America 27

scientificName: Brachyopa caesariata Moran and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: British Columbia; locality: Hixon;
decimalLatitude: 53.420261; decimalLongitude: -122.585961; eventDate: 06/23/1966;
year: 1966; month: 6; day: 23; verbatimEventDate: 23.vi.1966; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC_Diptera37560; recordedBy: E.D.A. Dyer;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Alaska; locality: Rte. 3, mi. 27011 mi. S
Anderson; decimalLatitude: 64.165328; decimalLongitude: -149.287278;
samplingProtocol: Malaise trap; eventDate: 1984-06/08-22/11; startDayOfYear: 173;
endDayOfYear: 224; year: 1984; month: 6; day: 22; verbatimEventDate:
22.Vi.-11.vill.1984; habitat: Alder-Poplar-Spruce; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera37561; recordedBy: S. & J. Peck;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: St. Williams; Norfolk;
decimalLatitude: 42.666667; decimalLongitude: -80.4; eventDate: 05/19/1970; year:
1970; month: 5; day: 19; verbatimEventDate: 19.v.1970; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera37566; recordedBy: D.M. Wood;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Alberta; locality: Vallyview;
decimalLatitude: 55.068678; decimalLongitude: -117.269575; eventDate: 06/04/1961;
year: 1961; month: 6; day: 4; verbatimEventDate: 4.vi.1961; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: CNC_Diptera37580; recordedBy: A.R.
Brooks; identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Constance L. South
March; decimalLatitude: 45.401397; decimalLongitude: -75.979717; eventDate:
06/16/1965; year: 1965; month: 6; day: 16; verbatimEventDate: 16.vi.1965;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera37581;

28

ae.

af.

ag.

ah.

al.

Skevington J et al

recordedBy: J.R. Vockeroth; identifiedBy: K. Moran; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: One Sided Lake;
decimalLatitude: 49.05; decimalLongitude: -93.883333; eventDate: 06/20/1960; year:
1960; month: 6; day: 20; verbatimEventDate: 20.vi.1960; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: CNC_Diptera37582; recordedBy: S.M. Clark;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Port Severn, 3 milles]
N.; decimalLatitude: 44.8; decimalLongitude: -79.716667; eventDate: 05/27/1959;
year: 1959; month: 5; day: 27; verbatimEventDate: 27.v.1959; habitat: black spruce
bog; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera37583; recordedBy: J.G. Chillcott; identifiedBy: K. Moran; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: Maryland; locality: Laurel; decimalLatitude:
39.083333; decimalLongitude: -76.833333; samplingProtocol: Malaise trap; eventDate:
05/25/1965; year: 1965; month: 5; day: 25; verbatimEventDate: 25.v.1965;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera37585;
identifiedBy: K. Moran; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: British Columbia; locality: 14 km East of
Coal River; decimalLatitude: 59.649875; decimalLongitude: -126.683892; eventDate:
1984-06/09-14/03; startDayOfYear: 166; endDayOfYear: 247; year: 1984; month: 6;
day: 14; verbatimEventDate: 14.vi.-3.ix.1984; habitat: Picea-Alnus forest;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera60558;
recordedBy: S. & J. Peck; identifiedBy: K. Moran; language: en; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: British Columbia; locality: 14 km East of
Coal River; decimalLatitude: 59.649875; decimalLongitude: -126.683892; eventDate:
1984-06/09-14/03; startDayOfYear: 166; endDayOfYear: 247; year: 1984; month: 6;

aj.

ak.

al.

am.

New Syrphidae (Diptera) of North-eastern North America 29

day: 14; verbatimEventDate: 14.vi.-3.ix.1984; habitat: Picea-Alnus forest;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera60559;
recordedBy: S. & J. Peck; identifiedBy: K. Moran; language: en; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: U.S.A.; stateProvince: New Hampshire; locality: Lancaster, Mount
Prospect; minimumElevationInMeters: 625; decimalLatitude: 44.449317;
decimalLongitude: -71.570889; eventDate: 06/19/1982; year: 1982; month: 6; day: 19;
verbatimEventDate: 19.vi.1982; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC_Diptera94649; recordedBy: J.R. Vockeroth; identifiedBy: K.
Moran; language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Manitoba; locality: 2 miles North East of
Treebank along Souris River; decimalLatitude: 49.666667; decimalLongitude:
-99.633333; eventDate: 06/09/1993; year: 1993; month: 6; day: 9; verbatimEventDate:
9.vi.1993; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera95516; recordedBy: B. Gallaway; identifiedBy: K. Moran; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Manitoba; locality: 2 miles North East of
Treebank along Souris River; decimalLatitude: 49.666667; decimalLongitude:
-99.633333; eventDate: 06/09/1993; year: 1993; month: 6; day: 9; verbatimEventDate:
9.vi.1993; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera95517; recordedBy: B. Gallaway; identifiedBy: K. Moran; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Brachyopa caesariata Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Plain-winged Sapeater;
genus: Brachyopa; specificEpithet: caesariata; scientificNameAuthorship: Moran and
Skevington; country: Canada; stateProvince: Ontario; locality: Orleans, Chapel Hill;
decimalLatitude: 45.466667; decimalLongitude: -75.516667; samplingProtocol:
sweeping; eventDate: 06/21/1995; year: 1995; month: 6; day: 21; verbatimEventDate:
21.vi.1995; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC_Diptera96849; recordedBy: J.R. Vockeroth; identifiedBy: K. Moran; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

Description

Size: Body length 5.2 to 7.9 mm; wing length 4.7 to 6.2 mm

30

Skevington J et al

Male: Head: Yellow to orange; face concave, covered in white pollen and bare, lower
face projecting forwards along the oral margin; gena pale pilose, shiny anteriorly and
gold pollinose posteriorly; frontal triangle shiny, bare except for gold pollinosity narrowly
along eye margin; vertex yellow, bare and gold pollinose, except ocellar triangle pale
pilose; occiput gold pollinose, pale pilose with some short black setae on dorsal %4;
antenna wholly yellow to orange, scape and pedicel with short, pale pile, pedicel about
as long as scape, postpedicel oval, about as long as wide, with distinct sensory pit on
inner surface, arista pubescent but not plumose; eye bare, broadly holoptic.

Thorax: Mesonotum yellow to orange, pale, black or mixed pale and black pilose and
gold pollinose except for two medial, bare stripes starting at anterior edge and running
about three-fourths the length of mesonotum and four bare vittae, two before
transverse suture and two after; scutellum yellow to orange, pale, black or mixed pale
and black pilose, with macrosetae; subscutellar fringe absent; postpronotum yellow to
orange, pale pilose and white pollinose; postalar callus yellow, pale pilose and shiny;
pleuron yellow, sparsely white pollinose; anterior anepisternum, katepimeron and
meron bare; posterior anepisternum and anepimeron pale pilose; katepisternum bare
on dorsal half; metasternum yellow, sparsely white pollinose, bare; legs yellow, except
Ath and 5th pro- and mesotarsomeres, as well as metatibia and tarsomeres black;
meso- and metacoxae sparsely white pollinose; femora pale pilose except meso- and
metafemur with black setae ventrally; tibia and tarsi mixed pale and black pilose; wing
entirely densely microtrichose; halter yellow; calypter white.

Abdomen: Oval, yellow to brown, pale pilose except tergites 2-4 black pilose medially,
shiny.

Genitalia: Epandrium subquadrate, longer than wide, narrows slightly dorsally; cercus
with laterally compressed, oval sclerotised outer portion covered in long pile and
membranous inner portion; surstylus short, with well separated dorsal and ventral
lobes, dorsal lobe short, ventrally compressed, ventral lobe large, laterally compressed,
rounded and elongate, dorsal lobe densely setose on posterior half of inner surface
and the posterior rim of the outer surface, ventral lobe with sparse row of pile on
posterior rim; subepandrial sclerite divided with one arm off of each surstylus, joined
only where it articulates with hypandrium; ejaculatory apodeme tiny, asymmetrical with
distal end slightly broadened; sperm pump inside the proximal end of the hyprandrium;
hypandrium narrow and elongated, rounded basally, ventrally with grooves
mediobasally; postgonite rectangular; phallus stubby and short.

Female: Similar to male except completely golden pollinose on frons between eyes in
addition to normal sexual dimorphism.

See Fig. 5.

New Syrphidae (Diptera) of North-eastern North America

Figure 5.

Brachyopa caesariata sp. nov.

a: Dorsal habitus, CNC_Diptera243273 ETSI
: Dorsal scutellum, CNC_Diptera243273 EES
c: Lateral habitus, CNC_Diptera243273 EES
d: Arista, CNC_Diptera243273 EE]
e
f

iy

: Lateral male genitalia, CNC_Diptera37549; scale bar 200 um EES
: Ventral male genitalia, CNC_Diptera37549; scale bar 200 um ETS]

31

32

Skevington J et al

Diagnosis

Katepisternum usually bare on dorsal half, rarely with 1-2 hairs. Mesonotum yellow in
colour. Never with wing spots. Arista pubescent. Abdomen and legs extensively yellow.

Etymology

The name is from the Latin caesariatus, meaning covered with hair, with reference to
the pilose arista.

Distribution

This species is found from Nova Scotia to British Columbia, north to Alaska and south
to Utah in the west and Maryland in the east.

Ecology

This uncommon species can be found flying from mid-May to late July but may be
found as early as the beginning of March in the southwest. Specimens have been
collected from Heracleum and Rubus plants and from black spruce bog and Alder-
Poplar-Spruce forest habitats.

Taxon discussion

This species is part of the B. punctipennis Curran 1925 complex of five species.
Skevington et al. (2019) incorrectly states that the name for this group is the B.
perplexa Curran 1922 complex. Brachyopa punctipennis s.s. is restricted to the
Western US as are three additional undescribed species in the complex. Like other
species in this complex, Brachyopa caesariata is found in the Western US; however it
is the only species whose range also extends to the Eastern US. Data for three of the
five undescribed species in this complex are available in BOLD. Brachyopa
punctipennis s.s. has not yet been sequenced. DNA sequences available for B.
caesariata are listed in (Table 2).

Common Name

The common name given to the species by Skevington et al. (2019) is Plain-winged
Sapeater.

New Syrphidae (Diptera) of North-eastern North America 33

Brachyopa cummingi Moran and Skevington, sp. n.

° ZooBank urn:lsid:zoobank.org:act:FE5C29DE-53A5-4CAD-B31A-039112D8DE28

Nomenclature
Brachyopa undescribed species 17-5 Skevington et al. (2019), page 216

Material

Holotype:

a. scientificName: Brachyopa cummingi Moran and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Somber Sapeater; genus:
Brachyopa; specificEpithet: cumming!; scientificNameAuthorship: Moran and
Skevington 2019; country: Canada; stateProvince: Ontario; locality: Ottawa;
decimalLatitude: 45.266667; decimalLongitude: -75.75; samplingProtocol: hand
collected; eventDate: 1986-05-14; year: 1986; month: 5; day: 14; verbatimEventDate:
14.v.1986; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC_Diptera106864; recordedBy: J.M. Cumming; identifiedBy: K. Moran; language:
en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

Description
Size: Body length 5.8 mm; wing length 5.4 mm

Male: Head: Yellow to orange; face concave, covered in white pollen and bare, lower
face projecting forwards along the oral margin; gena pale pilose, shiny anteriorly and
white pollinose posteriorly; frontal triangle shiny, bare except for white pollinosity and
short pale pile narrowly along eye margin; vertex yellow, except ocellar triangle black,
white pollinose and pale pilose; occiput white pollinose, pale pilose with some short
black setae on dorsal %; antenna wholly yellow to orange, scape and pedicel with
short, pale pile, pedicel about as long as scape, postpedicel oval, about twice as long
as wide, with minute sensory pit on inner surface, arista minutely pubescent; eye bare,
very narrowly dichoptic.

Thorax: Mesonotum dull brown, pale pilose, white pollinose except for two medial,
bare stripes starting at anterior edge and running about three-fourths the length of
mesonotum and four bare vittae, two before transverse suture and two after; scutellum
yellow, pale pilose, sparsely white pollinose and without macrosetae; subscutellar
fringe absent; postpronotum yellow, pale pilose and white pollinose; postalar callus
yellow, pale pilose and white pollinose; pleuron yellow, covered in white pollen; anterior
anepisternum, katepimeron and meron bare; posterior anepisternum and anepimeron
pale pilose; katepisternum discontinuously pale pilose; metasternum yellow, white
pollinose, bare; legs yellow, pale pilose; meso- and metacoxa sparsely white pollinose;
metafemur with black setulae along ventral side; wing entirely densely microtrichose;
halter yellow; calypter white.

34

Skevington J et al

Abdomen: Oval, unicolorous light brown, pale pilose, sparsely white pollinose.

Genitalia: Epandrium subquadrate, longer than wide, narrows slightly dorsally; cercus
with laterally compressed, subtriangular sclerotised outer portion covered in long pile
and membranous inner portion; surstylus short, with broadly fused, for ’/s the length of
the ventral lobe, dorsal and ventral lobes, dorsal lobe short, rounded apically, ventral
lobe large and elongate rectangularly, dorsal lobe densely setose on apex and with
dense cluster of long pile on outer surface just prior to apex, ventral lobe with sparse
smattering of pile on outer surface; subepandrial sclerite divided with one arm off of
each surstylus, joined only where it articulates with hypandrium; ejaculatory apodeme
narrow, straight; phallapodeme long and narrow; postgonite hooked ventrally, adjacent
to hypandrium; dorsal hypandrium with paired, pointed, long arms projecting from distal
end, with triangular tooth near tip; phallus simple tube within hypandrium.

Female: Unknown.

See Fig. 6.

Diagnosis

Arista pubescent but not plumose. Male postpedicel with distinct sensory pit. White
pollinose on scutum. Postpronotum and scutellum yellow in colour. Wing hyaline.
Katepisternum pilose on dorsal half. Scutellum pale pilose; completely pollinose.
Abdomen unicolorous light brown; completely pollinose.

Etymology

This species is named for the collector of the only known specimen. Jeff Cumming
works at the Canadian National Collection of Insects, Arachnids and Nematodes where
he specialises on Empidoidea systematics.

Distribution

The only known specimen of this species was collected in Ottawa, Ontario, Canada.

Ecology

This extremely rare fly has been collected once in mid-May (14 May 1986).

New Syrphidae (Diptera) of North-eastern North America

q . i. 3 i a
eo ee ee es ee

: “

Figure 6.

Brachyopa cummingi sp. nov.

a: Dorsal habitus, Holotype, CNC_Diptera106864 ETS]

b: Lateral habitus, Holotype, CNC_Diptera106864 EES]

c: Lateral, male genitalia, Holotype, CNC_Diptera106864 ETS]
d: Ventral, male genitalia, Holotype, CNC_Diptera106864 EES]
e: Antenna, Holotype, CNC_Diptera106864 ETS]

35

36

Skevington J et al

Taxon discussion

We were unable to sequence this specimen. Morphologically, it appears to be closely
related to B. daeckei Johnson 1917. Brachyopa daeckei lacks the distinct sensory pit
on the postpedicel that is present in B. cummingi. Additionally, B. daeckei has dark
brown instead of pale yellow postpronotum and scutelllum, gold instead of white
pollinosity on scutum and a pitch black instead of pale brown abdomen. Genitalia of B.
cummingi and B. daeckei are similar, except surstylar lobes of B. daecke/ are fused
over % the width of ventral lobe in comparison to B. cummingi in which they are fused
for 4 the width.

Common Name

The common name given to the species by Skevington et al. (2019) is Somber
Sapeater.

Hammerschmidtia sedmani Vockeroth, Moran and Skevington, sp. n.

ZooBank urn:|sid:zoobank.org:act:64C74904-E2A0-4EC1-8E42-0A18942BBA7C

Nomenclature

Brachyopa (Hammerschmidtia) sedmani Vockeroth manuscript name, in Thompson
(1991)

Hammerschmidtia undescribed species 1 , Skevington et al. (2019), page 218

Materials

Holotype:

a. scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
StateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322978; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298236;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

Paratypes:

a. scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
StateProvince: Ontario; locality: Algonquin Provincial Park, Opeongo Road;

New Syrphidae (Diptera) of North-eastern North America 37

decimalLatitude: 45.625911; decimalLongitude: -78.350753; eventDate: 2019-06-03;
year: 2018; month: 6; day: 3; verbatimEventDate: 3.vi.2018; habitat: on flowers of
Prunus virginiana; individualCount: 1; lifeStage: adult; catalogNumber: CNC1038609;
recordedBy: J.H.Skevington, M.M. Locke; identifiedBy: J.H. Skevington; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322976; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298237;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322976; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298238;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322976; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298239;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322976; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298240;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

38

Skevington J et al

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322976; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298241;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322976; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298242;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322978; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298243;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Base of Mount Rigaud; decimalLatitude: 45.466261;
decimalLongitude: -74.322978; eventDate: 2014-06-08; year: 2014; month: 6; day: 8;
verbatimEventDate: 8.vi.2014; habitat: on large fallen aspen log (on ground for about
one year); individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC298244;
recordedBy: J.H. Skevington; identifiedBy: J.H. Skevington; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Smooth Rock Falls, 56 km Northwest of Cochrane;

New Syrphidae (Diptera) of North-eastern North America 39

decimalLatitude: 49.275469; decimalLongitude: -81.629928; eventDate: 1984-06-03;
year: 1984; month: 6; day: 3; verbatimEventDate: 3.vi.1984; habitat: Picea-betula
forest; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera111405; recordedBy: S. & J. Peck; identifiedBy: J.R. Vockeroth;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Ottawa, Marlborough Forest Rideau Trail;
decimalLatitude: 45.05; decimalLongitude: -75.816667; eventDate: 2011-05-26; year:
2011; month: 5; day: 26; verbatimEventDate: 26.v.2011; habitat: near Rogers Pond;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera243287;
recordedBy: B.J. Sinclair; identifiedBy: J.R. Vockeroth; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: British Columbia; locality: Hixon; decimalLatitude: 53.4;
decimalLongitude: -122.566667; eventDate: 1966-06-07; year: 1966; month: 6; day: 7;
verbatimEventDate: 7.vi.1966; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera35319; recordedBy: E.D.A. Dyer; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Alberta; locality: Kananaskis, Forest Experimental Station Seebe;
decimalLatitude: 51.100879; decimalLongitude: -115.087692; eventDate: 1968-07-03;
year: 1968; month: 7; day: 3; verbatimEventDate: 3.vii.1968; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC_Diptera49254; recordedBy: H.J. Teskey;
identifiedBy: J.R. Vockeroth; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Ogoki; decimalLatitude: 51.633716; decimalLongitude:
-85.933333; eventDate: 1952-07-05; year: 1952; month: 7; day: 5; verbatimEventDate:
5.vil.1952; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC_Diptera49255; recordedBy: J.B. Wallis; identifiedBy: J.R. Vockeroth;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:

40

Skevington J et al

Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Laniel; decimalLatitude: 47.033333; decimalLongitude:
-79.266667; eventDate: 1931-06-10; year: 1931; month: 6; day: 10;
verbatimEventDate: 10.vi.1931; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera49256; recordedBy: H.S. Fleming; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Parke Reserve, Kamouraska County; decimalLatitude:
47.523387; decimalLongitude: -69.624739; eventDate: 1957-07-10; year: 1957; month:
7; day: 10; verbatimEventDate: 1957-07-10; individualCount: 1; sex: male; lifeStage:
adult; catalogNumber: CNC_Diptera49257; recordedBy: G.E. Shewell; identifiedBy:
J.R. Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: British Columbia; locality: Shames, 18 Miles South West of Terrace;
minimumElevation|InMeters: 32; decimalLatitude: 54.409648; decimalLongitude:
-128.935301; eventDate: 1960-07-17; year: 1960; month: 7; day: 17;
verbatimEventDate: 17.vil.1960; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera49258; recordedBy: B. Heming; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Mégantic; decimalLatitude: 45.583333;
decimalLongitude: -70.883333; eventDate: 1923-06-21; year: 1923; month: 6; day: 21;
verbatimEventDate: 21.vi.1923; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera49259; recordedBy: C.H. Curran; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Alberta; locality: Kananaskis, Forest Experimental Station Seebe;
decimalLatitude: 51.100883; decimalLongitude: -115.087692; eventDate: 1968-07-03;
year: 1968; month: 7; day: 3; verbatimEventDate: 3.vii.1968; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC_Diptera49260; recordedBy: H.J. Teskey;
identifiedBy: J.R. Vockeroth; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

New Syrphidae (Diptera) of North-eastern North America 41

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Harrington Lake, Gatineau Park [Lac Mousseau];
decimalLatitude: 45.568; decimalLongitude: -75.954; eventDate: 1954-06-09; year:
1954; month: 6; day: 9; verbatimEventDate: 9.vi.1954; individualCount: 1; sex: female;
lifeStage: adult; catalogNumber: CNC_Diptera49261; recordedBy: W.R. Richards;
identifiedBy: J.R. Vockeroth; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Masham Township; Gatineau; decimalLatitude:
45.683333; decimalLongitude: -76.05; eventDate: 1974-06-25; year: 1974; month: 6;
day: 25; verbatimEventDate: 25.vi.1974; habitat: sohagnum bog; individualCount: 1;
sex: female; lifeStage: adult; catalogNumber: CNC_Diptera49262; recordedBy: D.M.
Wood; identifiedBy: J.R. Vockeroth; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Masham Township; Gatineau; decimalLatitude:
45.683333; decimalLongitude: -76.05; eventDate: 1974-06-25; year: 1974; month: 6;
day: 25; verbatimEventDate: 25.vi.1974; habitat: sohagnum bog; individualCount: 1;
sex: female; lifeStage: adult; catalogNumber: CNC_Diptera49263; recordedBy: D.M.
Wood; identifiedBy: J.R. Vockeroth; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: British Columbia; locality: Barkerville; decimalLatitude: 53.083426;
decimalLongitude: -121.510871; eventDate: 1948-07-17; year: 1948; month: 7; day:
17; verbatimEventDate: 17.vii.1948; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC_Diptera49264; recordedBy: E.R. Buckell; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Mégantic; decimalLatitude: 45.583333;
decimalLongitude: -70.883333; eventDate: 1923-06-26; year: 1923; month: 6; day: 26;

42

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ab.

ac.

Skevington J et al

verbatimEventDate: 26.vi.1923; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC_Diptera49265; recordedBy: C.H. Curran; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: VockerVockeroth, Moran and Skevington 2019; country:
Canada; stateProvince: Ontario; locality: Ottawa; decimalLatitude: 45.423;
decimalLongitude: -75.698; eventDate: 1927-06-02; year: 1927; month: 6; day: 2;
verbatimEventDate: 2.vi.1927; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC_Diptera49266; recordedBy: C.H. Curran; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Duncan Lake, near Rupert; decimalLatitude:
45.681389; decimalLongitude: -76.050278; eventDate: 1971-06-17; year: 1971; month:
6; day: 17; verbatimEventDate: 17.vi.1971; individualCount: 1; sex: female; lifeStage:
adult; catalogNumber: CNC_Diptera49267; recordedBy: J.F. McAlpine; identifiedBy:
J.R. Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Orleans, Chapel Hill; decimalLatitude: 45.466667;
decimalLongitude: -75.516667; samplingProtocol: sweeping; eventDate: 1997-06-26;
year: 1997; month: 6; day: 26; verbatimEventDate: 26.vi.1997; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: CNC_Diptera49268; recordedBy: J.R.
Vockeroth; identifiedBy: J.R. Vockeroth; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Alberta; locality: Waterton Lakes National Park;
minimumElevationInMeters: 1300; decimalLatitude: 49.083313; decimalLongitude:
-113.916732; eventDate: 1980-06-21; year: 1980; month: 6; day: 21;
verbatimEventDate: 21.vi.1980; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC_Diptera49269; recordedBy: W.R.M. Mason; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;

ad.

ae.

af.

ag.

New Syrphidae (Diptera) of North-eastern North America 43

stateProvince: Ontario; locality: Orleans, Chapel Hill; decimalLatitude: 45.466667;
decimalLongitude: -75.516667; samplingProtocol: sweeping; eventDate: 1997-06-23;
year: 1997; month: 6; day: 23; verbatimEventDate: 23.vi.1997; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: CNC_Diptera49270; recordedBy: J.R.
Vockeroth; identifiedBy: J.R. Vockeroth; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: British Columbia; locality: Barkerville; decimalLatitude: 53.066667;
decimalLongitude: -121.516667; eventDate: 1948-07-17; year: 1948; month: 7; day:
17; verbatimEventDate: 17.vii.1948; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC_Diptera556; recordedBy: E.R. Buckell; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: One Sided Lake; decimalLatitude: 49.05;
decimalLongitude: -93.883333; eventDate: 1960-06-14; year: 1960; month: 6; day: 14;
verbatimEventDate: 14.vi.1960; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC_Diptera574; recordedBy: Kelton & Whitney; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Alberta; locality: Demmitt, 40 km North West of Beaverlodge;
decimalLatitude: 55.450035; decimalLongitude: -119.89992; samplingProtocol:
malaise trap; eventDate: 1984-06/09-12/06; eventTime: 164; startDayOfYear: 250;
year: 1984; month: 6; day: 12; verbatimEventDate: 12.vi.-6.xi.1984; habitat: Forest
Pupulus; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera60262; recordedBy: S. & J. Peck; identifiedBy: J.R. Vockeroth;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: British Columbia; locality: 14 km East of Coal River; decimalLatitude:
59.649875; decimalLongitude: -126.683892; eventDate: 1984-06/09-14/03; eveniTime:
166; startDayOfYear: 247; year: 1984; month: 6; day: 14; verbatimEventDate:
14.vi.-3.ix.1984; habitat: Picea-Alnus forest; individualCount: 1; sex: female; lifeStage:
adult; catalogNumber: CNC_Diptera60560; recordedBy: S. & J. Peck; identifiedBy: J.R.
Vockeroth; institutionCode: CNC; basisOfRecord: PreservedSpecimen

44

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al.

aj.

ak.

al.

Skevington J et al

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Alberta; locality: Demmitt, 40 km North West of Beaverlodge;
decimalLatitude: 55.450036; decimalLongitude: -119.899922; eventDate:
1984-06/09-12/06; eventTime: 164; startDayOfYear: 250; year: 1984; month: 6; day:
12; verbatimEventDate: 12.vi.-6.xi.1984; habitat: Forest; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: CNC_Diptera94610; recordedBy: S. & J.
Peck; identifiedBy: J.R. Vockeroth; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomencliaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Stittsville, Carleton County; decimalLatitude:
45.263479; decimalLongitude: -75.925163; eventDate: 1977-05-17/20; eventTime:
137; startDayOfYear: 140; year: 1977; month: 5; day: 17; verbatimEventDate:
17-20.v.1977; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera96475; recordedBy: M. Sanborne; identifiedBy: J.R. Vockeroth;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Stittsville, Carleton County; decimalLatitude:
45.263479; decimalLongitude: -75.925163; eventDate: 1977-05-20/22; eventTime:
140; startDayOfYear: 142; year: 1977; month: 5; day: 20; verbatimEventDate:
20-22.v.1977; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC_Diptera96476; recordedBy: M. Sanborne; identifiedBy: J.R. Vockeroth;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Orleans, Chapel Hill; decimalLatitude: 45.466667;
decimalLongitude: -75.516667; samplingProtocol: sweeping; eventDate: 1995-06-21;
year: 1995; month: 6; day: 21; verbatimEventDate: 21.vi.1995; habitat: meadow area;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera96844;
recordedBy: J.R. Vockeroth; identifiedBy: J.R. Vockeroth; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;

am.

an.

ao.

ap.

New Syrphidae (Diptera) of North-eastern North America 45

stateProvince: Ontario; locality: Orleans, Chapel Hill; decimalLatitude: 45.466667;
decimalLongitude: -75.516667; samplingProtocol: sweeping; eventDate: 1995-06-21;
year: 1995; month: 6; day: 21; verbatimEventDate: 21.vi.1995; habitat: meadow area;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera96845;
recordedBy: J.R. Vockeroth; identifiedBy: J.R. Vockeroth; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Orleans, Chapel Hill; decimalLatitude: 45.466667;
decimalLongitude: -75.516667; samplingProtocol: sweeping; eventDate: 1995-06-21;
year: 1995; month: 6; day: 21; verbatimEventDate: 21.vi.1995; habitat: meadow area;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera96846;
recordedBy: J.R. Vockeroth; identifiedBy: J.R. Vockeroth; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Ontario; locality: Orleans, Chapel Hill; decimalLatitude: 45.466667;
decimalLongitude: -75.516667; samplingProtocol: sweeping; eventDate: 1995-06-24:
year: 1995; month: 6; day: 24; verbatimEventDate: 24.vi.1995; habitat: meadow area;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera96847;
recordedBy: J.R. Vockeroth; identifiedBy: J.R. Vockeroth; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: New Brunswick; locality: Northumberland County, Nepisiguit Protected
Natural Area, 1.0km northwest of Popple Depot; decimalLatitude: 47.4024;
decimalLongitude: -66.522066; eventDate: 2017-06-22; year: 2017; month: 6; day: 22;
verbatimEventDate: 22.vi.2017; habitat: open Picea forest; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: JK10466; recordedBy: J. Klymko; identifiedBy:
J. Klymko; institutionCode: NBMB; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: New Brunswick; locality: Northumberland County, Nepisiguit Protected
Natural Area, 1.0km northwest of Popple Depot; decimalLatitude: 47.4024;
decimalLongitude: -66.522066; eventDate: 2017-06-22; year: 2017; month: 6; day: 22;
verbatimEventDate: 22.vi.2017; habitat: open Picea forest; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: JK10468; recordedBy: J. Klymko;
identifiedBy: J. Klymko; institutionCode: NBMB; basisOfRecord: PreservedSpecimen

46

aq.

ar.

as.

at.

au.

Skevington J et al

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: New Brunswick; locality: Northumberland County, Nepisiguit Protected
Natural Area, 1.0km northwest of Popple Depot; decimalLatitude: 47.4024;
decimalLongitude: -66.522066; eventDate: 2017-06-22; year: 2017; month: 6; day: 22;
verbatimEventDate: 22.vi.2017; habitat: open Picea forest; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: JK10470; recordedBy: J. Klymko; identifiedBy:
J. Klymko; institutionCode: NBMB; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: New Brunswick; locality: Northumberland County, Nepisiguit Protected
Natural Area, 1.0km northwest of Popple Depot; decimalLatitude: 47.4024;
decimalLongitude: -66.522066; eventDate: 2017-06-22; year: 2017; month: 6; day: 22;
verbatimEventDate: 22.vi.2017; habitat: open Picea forest; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: JK10474; recordedBy: J. Klymko; identifiedBy:
J. Klymko; institutionCode: NBMB; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: New Brunswick; locality: Restigouche Co., Jacquet R Gg PNA, woods
rd in mixed woods along Jacquet R.; decimalLatitude: 47.7733; decimalLongitude:
-66.1253; eventDate: 2010-06-25; year: 2010; month: 6; day: 25; verbatimEventDate:
25.vi.2010; habitat: Mixed Woods; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: JK736; recordedBy: J.Klymko,S.L.Robinson; identifiedBy: J. Klymko;
institutionCode: NBMB; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: New Hampshire; locality: Pittsburg, Rt. 3, Connecticut Lakes;
decimalLatitude: 45.104264; decimalLongitude: -71.259207; eventDate:
1974-06-22/24; eventTime: 142; startDayOfYear: 144; year: 1974; month: 6; day: 22;
verbatimEventDate: 22-24.vi.1974; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: USNMENT22534; recordedBy: B.J. & F.C. Thompson; identifiedBy:
F.C. Thompson; institutionCode: USNM; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: New Hampshire; locality: Pittsburg, Rt. 3, Connecticut Lakes;

av.

aw.

ax.

ay.

az.

New Syrphidae (Diptera) of North-eastern North America

decimalLatitude: 45.104264; decimalLongitude: -71.259207; eventDate:
1974-06-22/24; eventTime: 142; startDayOfYear: 144; year: 1974; month: 6; day: 22;
verbatimEventDate: 22-24.vi.1974; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: USNMENT22535; recordedBy: B.J. & F.C. Thompson; identifiedBy:
F.C. Thompson; institutionCode: USNM; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: New Hampshire; locality: Pittsburg, Rt. 3, Connecticut Lakes;
decimalLatitude: 45.104264; decimalLongitude: -71.259207; eventDate:
1974-06-22/24; eventTime: 142; startDayOfYear: 144; year: 1974; month: 6; day: 22;
verbatimEventDate: 22-24.vi.1974; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: USNMENT22536; recordedBy: B.J. & F.C. Thompson; identifiedBy:
F.C. Thompson; institutionCode: USNM; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: New Hampshire; locality: White Mountains, Dolly Copp;
decimalLatitude: 44.271; decimalLongitude: -71.305; eventDate: 1931-07-11; year:
1931; month: 7; day: 11; verbatimEventDate: 11.vii.1931; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: USNMENT249891; recordedBy: A.L.
Melander; identifiedBy: F.C. Thompson; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: British Columbia; locality: Kaslo; decimalLatitude: 49.907907;
decimalLongitude: -116.910358; eventDate: 06-15; month: 6; day: 15;
verbatimEventDate: 15.vi; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: USNMENT249892; recordedBy: R.P. Currie; identifiedBy: F.C.
Thompson; institutionCode: USNM; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;

47

stateProvince: Alberta; locality: Bilby; decimalLatitude: 53.683333; decimalLongitude:

-114.1; eventDate: 1924-06-11; year: 1924; month: 6; day: 11; verbatimEventDate:
11.vi.1924; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
USNMENT249893; recordedBy: O. Bryant; identifiedBy: F.C. Thompson;
institutionCode: USNM; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:

48

ba.

bb.

be.

bd.

Skevington J et al

Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: Michigan; locality: Macinac Co. Cedarville; decimalLatitude: 45.998;
decimalLongitude: -84.3656; eventDate: 1975-06-21; year: 1975; month: 6; day: 21;
verbatimEventDate: 21.vi.1975; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: USNMENT249894; recordedBy: D.D. Wilder; identifiedBy: F.C.
Thompson; institutionCode: CAS; basisOfRecord: PreservedSpecimen
scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington; country: Canada;
stateProvince: Quebec; locality: Duncan Lake, near Rupert; decimalLatitude:
45.681389; decimalLongitude: -76.050278; eventDate: 1996-06-29/30; eventTime:
181; startDayOfYear: 182; year: 1996; month: 6; day: 29; verbatimEventDate:
29-30.vi.1996; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC1185349; recordedBy: C. Kassebeer; identifiedBy: F.C. Thompson;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: Canada;
stateProvince: Quebec; locality: Duncan Lake, near Rupert; decimalLatitude:
45.681389; decimalLongitude: -76.050278; eventDate: 1996-06-29/30; eventTime:
181; startDayOfYear: 182; year: 1996; month: 6; day: 29; verbatimEventDate:
29-30.vi.1996; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC1185350; recordedBy: C. Kassebeer; identifiedBy: F.C. Thompson;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: New Hampshire; locality: Grafton Co., Bethlehem, Rt. 3, Rt. 3;
decimalLatitude: 44.281166; decimalLongitude: -71.687319; eventDate: 1973-06-18;
year: 1973; month: 6; day: 18; verbatimEventDate: 18.vi.1973; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: CNC1185351; recordedBy: B.J. & F.C.
Thompson; identifiedBy: F.C. Thompson; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: New Hampshire; locality: Jaffrey; decimalLatitude: 42.814629;
decimalLongitude: -72.023437; eventDate: 06-28; month: 6; day: 28;
verbatimEventDate: 28.vi; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC1185352; recordedBy: C.W. Johnson; identifiedBy: F.C.
Thompson; institutionCode: MCZ; basisOfRecord: PreservedSpecimen

New Syrphidae (Diptera) of North-eastern North America 49

be. scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: New York; locality: Essex Co., Keene Valley; decimalLatitude:
44.190122: decimalLongitude: -73.786586; eventDate: 1920-06-12; year: 1920; month:
6; day: 12; verbatimEventDate: 12.vi.1920; individualCount: 1; sex: female; lifeStage:
adult; catalogNumber: CNC1185353; recordedBy: H. Notman; identifiedBy: F.C.
Thompson; institutionCode: MCZ; basisOfRecord: PreservedSpecimen

bf. scientificName: Hammerschmidtia sedmani Vockeroth, Moran and Skevington;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Pale-bristled Logsitter; genus: Hammerschmidtia; specificEpithet: sedmani;
scientificNameAuthorship: Vockeroth, Moran and Skevington 2019; country: U.S.A.;
stateProvince: Pennsylvania; locality: Centre Co., Black Moshannon State Park;
decimalLatitude: 40.914568; decimalLongitude: -78.061262; eventDate: 1977-06-30;
year: 1977; month: 6; day: 30; verbatimEventDate: 30.vi.1977; individualCount: 1; sex:
female; lifeStage: adult; catalogNumber: CNC1185354; recordedBy: F.D. Fee;
identifiedBy: F.C. Thompson; institutionCode: ANSP; basisOfRecord:
PreservedSpecimen

Description
Size: Body length 7.5 to 9.5mm; wing length 6.0 to 7.7 mm

Male: Head: Orange; face with low tubercle, shiny medially, light (not obscuring orange
ground colour) white pollinose laterally; gena shiny anteriorly, white pollinose
posteriorly, white pilose; frontal triangle shiny, bare except for short white pile narrowly
along eye margin; front bare on ventral 1/3, yellow pilose dorsally; vertical triangle short
yellow pilose; occiput light white pollinose, white pilose with some short black setae on
dorsal %4; antenna yellow pilose; postpedicel oval, only slightly (1.2) longer than broad;
aristal pile long, with longest pili as long as pedicel width; eyes very narrowly dichoptic,
separated by width of an ommatidium; area of approximation short, about 5-6
ommatidia long.

Thorax: Dark brownish-orange; postpronotum yellow pilose except for a few black pili
medially; mesonotum shiny except light grey pollinose submedial vitta and notopleuron,
short black pilose except yellow pilose anteriorly and on notopleuron, with 3 supra-alar
and 1-2 postalar black bristles; scutellum shiny, black pilose, with broadly separated
apical black bristles and 1-3 subapical black bristles; pleuron sparsely grey pollinose,
white pilose except intermixed black pili on posterior anepisternum; anepistemum with
2-3 yellow posterodorsal bristles; katepistemum bare dorsally or with at most a few
short pili. Legs: pro- and mesocoxae orange, sparsely white pollinose, white pilose;
metacoxa dark brownish-orange, sparsely grey pollinose, yellow pilose, with 1-2 black
apicolateral bristles; trochanters orange, pale pilose; profemur orange, shiny, white
pilose except black pilose on posterior apical 12; mesofemur orange, black pilose
except white pilose on basoventral 12; metafemur brownish-orange, black pilose except

50

Skevington J et al

narrowly white pilose basally, with black setae on apico-ventral ’/3; pro- and mesotibiae
orange, black pilose; metatibia brownish-orange, black pilose, with a few black seta on
anterior side; tarsi orange except apical 2 tarsomeres brown, black pilose. Wing: halter
Orange; calypter white; wing microtrichose, hyaline except apical veins diffusely
margined with brown.

Abdomen: Brownish-orange, shiny, extensively white pilose, black pilose apicolaterally
on 2nd tergum and on apicomedial %% of 3rd and 4th terga.

Genitalia: Epandrium triangular, narrowing posteriorly; cercus with laterally
compressed, oval sclerotised outer portion covered in long pile and membranous inner
portion; surstylus long, fingerlike with rounded apex and with ventral tooth at about half
the length, densely setose on outer surface and loosely setose on inner surface;
subepandrial sclerite divided with one arm off of each surstylus, broad proximally and
joined only where it articulates with hypandrium; ejaculatory apodeme weakly fan-
shaped; phallapodeme long and narrow; postgonite small, quadrate; basiphallus claw-
like, broad; distiphallus simple tube.

Female: Face concave instead of tuberculate, eyes more broadly dichoptic.

See Fig. 7.

Diagnosis

Always orange with pale anepisternal bristles and a sparsely pilose katepisternum.

Etymology

This species is named for Yale Sedman in recognition of his early work on the male
genitalia and classification of brachyopine flower flies.

Distribution

This species is known from New Brunswick to British Columbia, south to Pennsylvania.

Ecology

This uncommon species is known to fly from mid-May to mid-July. They have been
found visiting Cornus canadensis, Heracleum, Physocarpus and Prunus virginiana and
are known from bogs and mixed woods. Several specimens were collected on a
recently fallen aspen log while they were searching for oviposition sites. Larvae of all
known species of Hammerschmidtia live in recently fallen aspens.

New Syrphidae (Diptera) of North-eastern North America

Figure 7.

Hammerschmidtia sedmani sp. nov.

a: Dorsal habitus, CNC_Diptera556 EES

b: Lateral habitus, CNC_Diptera574 Ei

c: Katepisternum, CNC_Diptera574 EE

d: Hind leg, CNC_Diptera574 ER]

e: Lateral of male genitalia, CNC_Diptera49259; scale bar 500 um EES]
f: Ventral of male genitalia, CNC_Diptera49259; scale bar 200 um EES

51

52 Skevington J et al

Taxon discussion

DNA sequences of this species are available from BOLD and GenBank (Table 2). This
new species is similar to H. rufa, but can be distinguished by the black anepisternal
bristles and katepisternum that is densely pilose.

Common Name

The common name given to the species by Skevington et al. (2019) is Pale-bristled
Logsitter.

Microdon (Microdon) scauros Skevington and Locke, sp. n.

e ZooBank urn:|sid:zoobank.org:act:0DDD7DE3-C6F7-490D-A0B3-AA870F1C4815

Nomenclature
Microdon undescribed species 17-1 Skevington et al. (2019), page 46

Materials

Holotype:

a. scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-03; year: 1986; month: 9; day: 3; habitat: open grassy old field;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44169; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

Paratypes:

a. scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Maryland; locality: Allegany County, Fifteen-Mile Creek, Green Ridge State Forest;
decimalLatitude: 39.617; decimalLongitude: -78.383; samplingProtocol: hand collected;
eventDate: 1978-07-06; year: 1978; month: 7; day: 6; individualCount: 1; sex: female;
lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen44200; recordedBy: F.D.
Fee; identifiedBy: Jeffrey H. Skevington; dateldentified: 2017; language: en;
institutionCode: ANSP; basisOfRecord: PreservedSpecimen

b. scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:

New Syrphidae (Diptera) of North-eastern North America 53

Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1987-08-30; year: 1987; month: 8; day: 30; habitat: perch/veg.,
dry weedy meadow; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44176; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1987-09-15; year: 1987; month: 9; day: 15; habitat: low veg.
adjacent to marsh; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44179; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1987-09-15; year: 1987; month: 9; day: 15; habitat: low veg.
adjacent to marsh; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44180; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1978-09-02; year: 1978; month: 9; day: 2; individualCount: 1;
sex: male; lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen44184;
recordedBy: F.D. Fee; identifiedBy: Jeffrey H. Skevington; dateldentified: 2017;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;

54

Skevington J et al

decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1978-09-02; year: 1978; month: 9; day: 2; individualCount: 1;
sex: male; lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen44185;
recordedBy: F.D. Fee; identifiedBy: Jeffrey H. Skevington; dateldentified: 2017;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1978-09-02; year: 1978; month: 9; day: 2; individualCount: 1;
sex: male; lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen44186;
recordedBy: F.D. Fee; identifiedBy: Jeffrey H. Skevington; dateldentified: 2017;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1978-09-02; year: 1978; month: 9; day: 2; individualCount: 1;
sex: male; lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen44187;
recordedBy: F.D. Fee; identifiedBy: Jeffrey H. Skevington; dateldentified: 2017;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1978-09-02; year: 1978; month: 9; day: 2; individualCount: 1;
sex: male; lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen44188;
recordedBy: F.D. Fee; identifiedBy: Jeffrey H. Skevington; dateldentified: 2017;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1978-09-02; year: 1978; month: 9; day: 2; habitat: veg., shrubby
meadow; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44199; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

New Syrphidae (Diptera) of North-eastern North America 55

scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Ridge & Nursery Roads, Barree Township;
decimalLatitude: 40.642501; decimalLongitude: -77.908807; samplingProtocol: hand
collected; eventDate: 1987-08-31; year: 1987; month: 8; day: 31; habitat: ex. low veg.,
dry weedy field; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen45193; recordedBy: F.D. Fee; otherCatalogNumbers:
CCDB-26861; identifiedBy: Jeffrey H. Skevington; dateldentified: 2017; language: en;
institutionCode: ANSP; basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Route 26, 1.1 km west of Whipple Dam
State Park; decimalLatitude: 40.69385; decimalLongitude: -77.857447;
samplingProtocol!: hand collected; eventDate: 1988-08-27; year: 1988; month: 8; day:
27; habitat: veg., sedge meadow; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: Jeff_Skevington_Specimen44181; recordedBy: F.D. Fee; identifiedBy:
Jeffrey H. Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Route 26, 1.1 km west of Whipple Dam
State Park; decimalLatitude: 40.69385; decimalLongitude: -77.857447;
samplingProtocol: hand collected; eventDate: 1988-08-27; year: 1988; month: 8; day:
27; habitat: veg., sedge meadow; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: Jeff_Skevington_Specimen44182; recordedBy: F.D. Fee; identifiedBy:
Jeffrey H. Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Stone Valley Recreational Area;
decimalLatitude: 40.659991; decimalLongitude: -77.919963; samplingProtocol: hand
collected; eventDate: 1987-08-30; year: 1987; month: 8; day: 30; habitat: veg., shrubby
meadow; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44177; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;

56

Skevington J et al

family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Stone Valley Recreational Area;
decimalLatitude: 40.659991; decimalLongitude: -77.919963; samplingProtocol: hand
collected; eventDate: 1987-08-30; year: 1987; month: 8; day: 30; habitat: veg., shrubby
meadow; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44178; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Stone Valley Recreational Area;
decimalLatitude: 40.659991; decimalLongitude: -77.919963; samplingProtocol: hand
collected; eventDate: 1987-08-31; year: 1987; month: 8; day: 31; habitat: veg., shrubby
meadow; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44197; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Stone Valley Recreational Area;
decimalLatitude: 40.659991; decimalLongitude: -77.919963; samplingProtocol: hand
collected; eventDate: 1987-08-31; year: 1987; month: 8; day: 31; habitat: veg., shrubby
meadow; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44198; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-08-31; year: 1986; month: 8; day: 31; habitat: ex: clearing among
white pines/old field; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen29871; recordedBy: F.D. Fee; otherCatalogNumbers:
CCDB-26861; identifiedBy: Jeffrey H. Skevington; date!ldentified: 2017; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:

New Syrphidae (Diptera) of North-eastern North America 57

Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-08-31; year: 1986; month: 8; day: 31; habitat: ex: clearing among
white pines/old field; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen29872; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-08-31; year: 1986; month: 8; day: 31; habitat: old field clearing/white
pines; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44167; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-08-31; year: 1986; month: 8; day: 31; habitat: old field clearing/white
pines; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44168; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-03; year: 1986; month: 9; day: 3; habitat: old field, clearing, white
pines; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44170; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:

58

aa.

Skevington J et al

40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-06; year: 1986; month: 9; day: 6; habitat: low veg,., old field;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44171; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-06; year: 1986; month: 9; day: 6; habitat: low veg,., old field;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44172; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-06; year: 1986; month: 9; day: 6; habitat: clearing-white pines-old
field; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44173; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-11; year: 1986; month: 9; day: 11; habitat: Perching-grass-old
field; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44174; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-13; year: 1986; month: 9; day: 13; habitat: clearing, old field;

ab.

ac.

ad.

ae.

New Syrphidae (Diptera) of North-eastern North America 59

individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44175; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: USNM;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1989-07-29; year: 1989; month: 7; day: 29; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen44183; recordedBy: F.D.
Fee; identifiedBy: Jeffrey H. Skevington; dateldentified: 2017; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-08-31; year: 1986; month: 8; day: 31; habitat: clearing, white pine/old
field; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44189; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-04; year: 1986; month: 9; day: 4; habitat: open grassy old field;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44190; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: USNM;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-06; year: 1986; month: 9; day: 6; habitat: low veg,., old field;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44191; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.

60

af.

ag.

ah.

al.

Skevington J et al

Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-06; year: 1986; month: 9; day: 6; habitat: low veg,., old field;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44192:; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-06; year: 1986; month: 9; day: 6; habitat: open grassy mowed
area; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44193; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-06; year: 1986; month: 9; day: 6; habitat: low veg,., old field;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44194; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-11; year: 1986; month: 9; day: 11; habitat: Perching-grasses-old
field; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44195; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

New Syrphidae (Diptera) of North-eastern North America 61

aj. scientificName: Microdon scauros Skevington and Locke; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntingdon County, Whipple Dam State Park; decimalLatitude:
40.69385; decimalLongitude: -77.857447; samplingProtocol: hand collected;
eventDate: 1986-09-13; year: 1986; month: 9; day: 13; habitat: clearing, old field;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44196; recordedBy: F.D. Fee; identifiedBy: Jeffrey H.
Skevington; dateldentified: 2017; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

ak. scientificName: Microdon scauros Skevington and Locke; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Big-footed Ant Fly; genus:
Microdon; subgenus: Microdon; specificEpithet: scauros; scientificNameAuthorship:
Skevington and Locke 2019; country: United States of America; stateProvince:
Pennsylvania; locality: Huntington County, Route 26, 1.1 km west of Whipple Dam
State Park; decimalLatitude: 40.684219; decimalLongitude: -77.863903;
samplingProtocol: hand collected; eventDate: 1988-09-01; year: 1988; month: 9; day:
1; habitat: ex. vegetation, dry weedy hillside; individualCount: 1; sex: female; lifeStage:
adult; catalogNumber: Jeff_Skevington_Specimen18258; recordedBy: F.D. Fee;
identifiedBy: Jeffrey H. Skevington; dateldentified: 2017; language: en; institutionCode:
ANSP; basisOfRecord: PreservedSpecimen

Description
Size: Body length 7.5 to 11.9 mm; wing length 4.7 to 7.2 mm

Male: Head: Brown, shining, entirely white pilose; occiput weakly grey pollinose, white
pilose; scape pale brown, pedicel and postpedicel contrasting dark brown; postpedicel
with a short sensory groove (about 0.1x length of postpedicel) below base of arista;
antennal ratio (Scape:pedicel:postpedicel) 4:1:7; eye bare; dichoptic, with eyes
separated by 0.6-0.7x length of scape.

Thorax: Brown, shining; scutum and scutellum entirely white pilose; scutellum with
prominent pair of apical, inward-pointing calcars; postpronotum white pilose; anterior
anepisternum mostly bare with white pile only adjacent to posterior anepisternum,
posterior anepisternum entirely white pilose, bare ventrally; katepisternum mostly
shining brown with white pile only on posterodorsal corner and in a small ventral patch
between fore and mid coxae; katepisternum bare and shining ventrally, pilose dorsally;
meron and katepimeron bare, shining brown; anepimeron and anatergum sparsely
pilose, dull; halter pale yellow; calypter pale yellow with white fringe; femora, coxae and
trochanters dark brown, white pilose; tibiae pale yellowish-brown with dark brown
medial band, white pilose; pro- and mesotarsi pale yellowish-brown with white pile
dorsally and shorter golden pile ventrally and laterally; metabasitarsis dark brown,
swollen and cylindrical; other metatarsi similar to those of pro- and mesolegs; wing light
brownish, microtrichose except for narrow bare area along leading edges of bm and br.

62

Skevington J et al

Abdomen: Entirely dark brown dorsally and white pilose; sternites and terminalia paler
brown, white pilose.

Genitalia: Epandrium compact, similar in length to surstylus; cercus subquadrate, with
apex rounded and slightly wider at base; surstylus broad base, finger-shaped distally,
with broad short spines ventroapically; phallus broad basally, ending in 2 stout tubules;
apical part of hypandrium narrow, leaf-like; ejaculatory apodeme flat, rounded.

Female: Similar to male, differing as follows: Antennal ratio 3-4:1:5-6; terminalia dark
brown except cerci paler.

See Fig. 8 & Fig. 9.

Diagnosis

Dark with prominent inward-pointing calcars on the scutellum, swollen metabasitarsus
particularly evident on the male, postpedicel longer than scape, wings shorter than M.
trisits and completely microtrichose (including cell bm).

Etymology

Based on the Greek word skauros, for having large and swollen ankles, with reference
to the swollen metabasitarsus of this species.

Distribution

Specimens are known from five sites in a small area of Pennsylvania and Maryland
and were all collected by Frank Fee.

Ecology

This species appears to be active late in the season and is known to fly from late
August to mid-September. One specimen was found flying earlier, in late July. This
species is known from a small area of the north-eastern United States and is
apparently rare. The known specimens were found in old fields and sedge meadows
with adjacent pine-dominated forests or marshland.

Taxon discussion

Most closely resembles Microdon tristis Loew, 1864. Microdon tristis can be
distinguished by prominent, outward-pointing calcars on the scutellum, metabasitarsus
only slightly expanded (not prominently swollen), orange tibiae contrast with dark
femora, postpedicel as long or longer than scape, wing with cell bm bare
posterobasally. DNA was obtained from a single specimen (Table 2). Although
morphologically most similar to M. tristis , it is most similar genetically to M. globosus

New Syrphidae (Diptera) of North-eastern North America 63

(Fabricius 1805). Microdon scauros and M. globosus, while genetically similar, are
quite different morphologically and clearly distinguished by Skevington et al. (2019).

Figure 8.

Microdon scauros, Sp. nov.

a: Dorsal habitus, Holotype, Jeff_Skevington_Specimen44169
b: Lateral habitus, Holotype, Jeff_Skevington_Specimen44169
c: Antennae, Holotype, Jeff_Skevington_Specimen44169

d: Scutellum, Jeff_Skevington_Specimen44200

e: Male hind tarsi, Holotype, Jeff_Skevington_ Specimen44169
f: Female hind tarsi, Jeff_Skevington_Specimen44200 [E&I

64

Skevington J et al

Figure 9.

Microdon scauros male genitalia, sp. nov.

a: Lateral of male genitalia, Jeff_Skevington_Specimen44181; scale bar 500 um EES
b: Ventral of male genitalia, Jeff_Skevington_Specimen44181; scale bar 500 um ERS]
c: Lateral of ejaculatory apodeme and sperm pump, Jeff_Skevington_Specimen44181;

scale bar 200 um EES
d: Lateral of phallus, hypandrium and related structures, Jeff_Skevington_Specimen44183,

scale bar 200 um EES

Common Name

The common name given to the species by Skevington et al. (2019) is Big-footed Ant
Fly.

New Syrphidae (Diptera) of North-eastern North America 65

Mixogaster fattigi Locke, Skevington and Greene, sp. n.

° ZooBank urn:lsid:zoobank.org:act:42F8F983-1C7B-49BD-B015-8C50A2CB6884

Nomenclature

Mixogaster fattigi Greene manuscript name (unpublished, undated manuscript)
Mixogaster fattig’ Weems manuscript name (Weems 1953), page 253
Mixogaster fattigi Greene manuscript name in Thompson (1991)

Mixogaster urania Hull manuscript name in Thompson (1991)

Mixogaster undescribed species 1 Skevington et al. (2019), page 56

Materials

Holotype:

a. scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster, specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Atlanta; decimalLatitude: 33.75; decimalLongitude: -84.383333; eventDate:
1941-07-08; year: 1941; month: 7; day: 8; verbatimEventDate: 8.vii.1941;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen17086; recordedBy: P.W. Fattig; identifiedBy: J.H.
Skevington; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

Paratypes:

a. scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Tennessee;
locality: Great Smoky Mountains National Park, Cades Cove, Parson's Branch Road;
decimalLatitude: 35.6; decimalLongitude: -83.85; samplingProtocol: Malaise trap;
eventDate: 1999-07-10; year: 1999; month: 7; day: 10; verbatimEventDate:
10-12.vii.1999; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC1129651; recordedBy: G. Steck and B. Sutton; identifiedBy: J.H. Skevington;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

b. scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Tennessee;
locality: Great Smoky Mountains National Park, Cades Cove, Parson\'s Branch Road;
decimalLatitude: 35.6; decimalLongitude: -83.85; samplingProtocol: Malaise trap;
eventDate: 1999-07-10/12; year: 1999; month: 7; day: 10; verbatimEventDate:

66

Skevington J et al

10-12.vii.1999; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC1129652; recordedBy: G. Steck and B. Sutton; identifiedBy: J.H. Skevington;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Tennessee;
locality: Great Smoky Mountains National Park, Cades Cove, Parson\'s Branch Road;
decimalLatitude: 35.6; decimalLongitude: -83.85; samplingProtocol: Malaise trap;
eventDate: 1999-07-10/12; year: 1999; month: 7; day: 10; verbatimEventDate:
10-12.vii.1999; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC1129653; recordedBy: G. Steck and B. Sutton; identifiedBy: J.H. Skevington;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Tennessee;
locality: Great Smoky Mountains National Park, Cades Cove, Parson\'s Branch Road;
decimalLatitude: 35.6; decimalLongitude: -83.85; samplingProtocol: Malaise trap;
eventDate: 1999-07-10/12; year: 1999; month: 7; day: 10; verbatimEventDate:
10-12.vii.1999; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC1129654; recordedBy: G. Steck and B. Sutton; identifiedBy: J.H. Skevington;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Louisiana;
locality: Bayou Chicot, Evangeline Parish; decimalLatitude: 30.815558;
decimalLongitude: -92.350645; eventDate: 1971-09-17; year: 1971; month: 8; day: 17;
verbatimEventDate: 17.vili.1971; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera102833; recordedBy: D.Shanek; identifiedBy: J.H.
Skevington; language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster, specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Louisiana;
locality: Bayou Chicot, Evangeline Parish; decimalLatitude: 30.815558;
decimalLongitude: -92.350645; eventDate: 1971-09-17; year: 1971; month: 8; day: 17;
verbatimEventDate: 17.vili.1971; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera102834; recordedBy: D.Shanek; identifiedBy: J.H.
Skevington; language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi; scientificNameAuthorship:
Locke, Skevington and Greene; country: U.S.A.; stateProvince: Georgia; locality:

New Syrphidae (Diptera) of North-eastern North America 67

Atlanta; decimalLatitude: 33.75; decimalLongitude: -84.383333; eventDate:
1941-06-26; year: 1941; month: 6; day: 26; verbatimEventDate: 26.vi.1941;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera92828;
recordedBy: P.W. Fattig; identifiedBy: J.H. Skevington; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Atlanta; decimalLatitude: 33.75; decimalLongitude: -84.383333; eventDate:
1942-06-30; year: 1942; month: 6; day: 30; verbatimEventDate: 30.vi.1942;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera92829;
recordedBy: P.W. Fattig; identifiedBy: J.H. Skevington; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster, specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Atlanta; decimalLatitude: 33.75; decimalLongitude: -84.383333; eventDate:
1942-07-15; year: 1942; month: 7; day: 15; verbatimEventDate: 15.vii.1942;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera92831;
recordedBy: P.W. Fattig; identifiedBy: J.H. Skevington; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Texas; locality:
Lamar Co., Camp Maxey; decimalLatitude: 33.667222; decimalLongitude: -95.566389;
samplingProtocol: Malaise trap; eventDate: 2003-07-01; year: 2003; month: 7; day: 1;
verbatimEventDate: 1-15.viil.2003; habitat: in pine forest; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen25513; recordedBy: W.
Godwin; otherCatalogNumbers: accession#SFASU lot 53; identifiedBy: J.H.
Skevington; language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Mississippi;
locality: Winston Co., 13 mi. S. Starkville; decimalLatitude: 33.286; decimalLongitude:
-88.91; samplingProtocol: Malaise trap; eventDate: 1982-06-28; year: 1982; month: 6;
day: 28; verbatimEventDate: 28.vi.1982; habitat: pine forest; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen45174; recordedBy:
N. Bedwell; identifiedBy: J.H. Skevington; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:

Skevington J et al

Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Florida; locality:
Alachua Co., San Felasco Hammock; decimalLatitude: 29.73; decimalLongitude:
-82.43; samplingProtocol: carbon dioxide baited flight interception trap; eventDate:
1977-06-02; year: 1977; month: 6; day: 2; verbatimEventDate: 2.vi.1977;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen45175; recordedBy: H. V. Weems, Jr. & G. B. Fairchild;
identifiedBy: J.H. Skevington; language: en; institutionCode: ANSP; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Mississippi;
locality: Oktibbeha Co., Dorman Lk.; decimalLatitude: 33.3394; decimalLongitude:
-88.8728; samplingProtocol: Malaise trap; eventDate: 1981-06-20/27; year: 1981;
month: 6; day: 20; verbatimEventDate: 20-27.vi.1981; habitat: hardwood forest;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: MEM92633;
recordedBy: R.L. Brown; identifiedBy: J.H. Skevington; language: en; institutionCode:
MEMU; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Mississippi;
locality: Oktibbeha Co., 6 mi. SW of Starkville; decimalLatitude: 33.3646;
decimalLongitude: -88.873; samplingProtocol: Malaise trap; eventDate: 1984-07-18/22;
year: 1984; month: 7; day: 18; verbatimEventDate: 18-22.viil.1984; habitat: mixed pine/
hardwood forest; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
MEM92634; recordedBy: R.L. & B.B. Brown; identifiedBy: J.H. Skevington; language:
en; institutionCode: MEMU; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster, specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Mississippi;
locality: Oktibbeha Co., 5 mi. S of Starkville; decimalLatitude: 33.379722:
decimalLongitude: -88.828889; samplingProtocol: Malaise trap; eventDate:
2000-06-20/26; year: 2000; month: 6; day: 20; verbatimEventDate: 20-26.vi.2000;
habitat: mixed pine/oak forest; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: MEM92716; recordedBy: R.L. Brown; identifiedBy: J.H. Skevington;
language: en; institutionCode: MEMU; basisOfRecord: PreservedSpecimen
scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster, specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Mississippi;
locality: Oktibbeha Co., 5 mi. S of Starkville; decimalLatitude: 33.379722;
decimalLongitude: -88.828889; samplingProtocol: Malaise trap; eventDate:
2000-06-13/19; year: 2000; month: 6; day: 13; verbatimEventDate: 13-19.vi.2000;

New Syrphidae (Diptera) of North-eastern North America 69

habitat: mixed pine/oak forest; individualCount: 2; sex: male; lifeStage: adult;
catalogNumber: MEM92717; recordedBy: R.L. Brown; identifiedBy: J.H. Skevington;
language: en; institutionCode: MEMU; basisOfRecord: PreservedSpecimen
scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Mississippi;
locality: Oktibbeha Co., T18N R14E Sec. 23; decimalLatitude: 33.43;
decimalLongitude: -88.88; eventDate: 1990-06-26; year: 1990; month: 6; day: 26;
verbatimEventDate: 26.vi.1990; habitat: mixed pine/oak forest; individualCount: 1; sex:
male; lifeStage: adult; catalogNumber: MEM92718; recordedBy: R.L. & B.B. Brown;
identifiedBy: J.H. Skevington; language: en; institutionCode: MEMU; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene; country: U.S.A.; stateProvince: Mississippi; !ocality:
Oktibbeha Co., 6 mi SW Starkville; decimalLatitude: 33.3646; decimalLongitude:
-88.873; samplingProtocol: Malaise trap; eventDate: 1990-07-11; year: 1990; month: 7;
day: 11; verbatimEventDate: 11.vii.1990; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: MEM92719; recordedBy: R.L. & B.B. Brown; identifiedBy: J.H.
Skevington; language: en; institutionCode: MEMU; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Mississippi;
locality: Oktibbeha Co., 6 mi SW Starkville; decimalLatitude: 33.3646;
decimalLongitude: -88.873; samplingProtocol: Malaise trap; eventDate: 1984-07-06;
year: 1984; month: 7; day: 6; verbatimEventDate: 6-7.vii.1984; individualCount: 2; sex:
male; lifeStage: adult; catalogNumber: MEM92720; recordedBy: R.L. & B.B. Brown;
identifiedBy: J.H. Skevington; language: en; institutionCode: MEMU; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Mississippi;
locality: Oktibbeha Co., 6 mi SW Starkville; decimalLatitude: 33.3646;
decimalLongitude: -88.873; samplingProtocol: Malaise trap; eventDate: 1984-07-18/22;
year: 1984; month: 7; day: 18; verbatimEventDate: 18-22.vii.1984; habitat: mixed pine/
hardwood forest; individualCount: 3; sex: male; lifeStage: adult; catalogNumber:
MEM92721; recordedBy: R.L. & B.B. Brown; identifiedBy: J.H. Skevington; language:
en; institutionCode: MEMU; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:

70

Skevington J et al

Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Virginia; locality:
Falls Church; decimalLatitude: 38.883333; decimalLongitude: -77.166667; eventDate:
1951-07-22; year: 1951; month: 7; day: 22; verbatimEventDate: 22.vii.1951; habitat:
Stream margin; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
USNMENT19965; recordedBy: W.W. Wirth; identifiedBy: J.H. Skevington; language:
en; institutionCode: USNM; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Atlanta; decimalLatitude: 33.75; decimalLongitude: -84.383333; eventDate:
1942-06-09; year: 1942; month: 7; day: 9; verbatimEventDate: 9.vii.1942;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: USNMENT247135;
recordedBy: P.W. Fattig; identifiedBy: J.H. Skevington; language: en; institutionCode:
USNM; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Atlanta; decimalLatitude: 33.75; decimalLongitude: -84.383333; eventDate:
1973-07-07; year: 1973; month: 7; day: 7; verbatimEventDate: 7.vii.1973;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: USNMENT247136;
recordedBy: H.D. Pratt; identifiedBy: J.H. Skevington; language: en; institutionCode:
USNM; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Clarke County, Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667;
samplingProtocol: Malaise trap; eventDate: 1976-10/12; year: 1976; month: 10;
verbatimEventDate: x.-xii.1976; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: USNMENT247137; recordedBy: R.W. Matthews; identifiedBy: J.H.
Skevington; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Clarke County, Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667;
samplingProtocol: Malaise trap; eventDate: 1976-10/12; year: 1976; month: 10;
verbatimEventDate: x.-xii.1976; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: USNMENT247138; recordedBy: R.W. Matthews; identifiedBy: J.H.
Skevington; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

aa.

ab.

ac.

ad.

New Syrphidae (Diptera) of North-eastern North America 71

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667; samplingProtocol:
Malaise trap; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
USNMENT 247139; recordedBy: R. Duffield; identifiedBy: J.H. Skevington; language:
en; institutionCode: USNM; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Clarke County; decimalLatitude: 33.95; decimalLongitude: -83.366667;
samplingProtocol: Malaise trap; eventDate: 1971-07-04/10; year: 1971; month: 7; day:
4; verbatinEventDate: 4-10.vil.1971; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: USNMENT247140; recordedBy: R.T. Franklin; identifiedBy: J.H.
Skevington; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667; samplingProtocol:
Malaise trap; eventDate: 1971-06-30; year: 1971; month: 6; day: 30;
verbatimEventDate: 30.vi.1971; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: USNMENT247141; recordedBy: R. Franklin; identifiedBy: J.H.
Skevington; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Clarke County, Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667;
eventDate: 1976-07; year: 1976; month: 7; verbatimEventDate: vil.1976;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: USNMENT247142;
identifiedBy: J.H. Skevington; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667; samplingProtocol:
Malaise trap; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
USNMENT 247143; recordedBy: R. Duffield; identifiedBy: J.H. Skevington; language:
en; institutionCode: USNM; basisOfRecord: PreservedSpecimen

72

ae.

af.

ag.

ah.

al.

Skevington J et al

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Bogart; decimalLatitude: 33.949258; decimalLongitude: -83.5346; eventDate:
1974-07-07; year: 1974; month: 7; day: 7; verbatimEventDate: 7.vii.1974;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: USNMENT247144;
recordedBy: A. Lavallee; identifiedBy: J.H. Skevington; language: en; institutionCode:
USNM; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Clarke County, Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667;
samplingProtocol: Malaise trap; eventDate: 1976-10/12; year: 1976; month: 10;
verbatimEventDate: x.-xii.1976; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: USNMENT247145; recordedBy: R.W. Matthews; identifiedBy: J.H.
Skevington; language: en; institutionCode: USNM; basisOfRecord:
PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Bogart; decimalLatitude: 33.949258; decimalLongitude: -83.5346; eventDate:
1972-07-07; year: 1972; month: 7; day: 7; verbatimEventDate: 7.vii.1972;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: USNMENT247146;
recordedBy: A. Lavallee; identifiedBy: J.H. Skevington; language: en; institutionCode:
USNM; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster, specificEpithet: fattigr, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667; samplingProtocol:
Malaise trap; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
USNMENT 247147; recordedBy: R. Duffield; identifiedBy: J.H. Skevington; language:
en; institutionCode: USNM; basisOfRecord: PreservedSpecimen

scientificName: Mixogaster fattigi Locke, Skevington and Greene;
nomenciaturalStatus: new species; kingdom: Animalia; phylum: Arthropoda; class:
Hexapoda; order: Diptera; family: Syrphidae; taxonRank: species; vernacularName:
Fattig's Ant Fly; genus: Mixogaster,; specificEpithet: fattigi, scientificNameAuthorship:
Locke, Skevington and Greene 2019; country: U.S.A.; stateProvince: Georgia; locality:
Athens; decimalLatitude: 33.95; decimalLongitude: -83.366667; samplingProtocol:
Malaise trap; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
USNMENT 247148; recordedBy: R. Duffield; identifiedBy: J.H. Skevington; language:
en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

New Syrphidae (Diptera) of North-eastern North America 73

Description
Size: Body length: 10.3-14.6 mm; wing length 7.4-9.1 mm

Male: Head: Face slightly convex, parallel-sided, projecting downwards slightly below
the eye, yellow, shining with yellow pile, sometimes with darkened area below
antennae rarely extending to black, narrow medial stripe not reaching the oral margin;
gena yellow, reduced, yellow pilose, separated from face by black stripe extending from
eye to oral margin; frons with broad black transverse band along lower edge, black
pilose laterally, some admixed yellow pile, shining medially; vertex yellow, black pilose;
ocellar triangle black with black extending posteriorly to a point; antenna elongate,
dark, scape and pedicel yellow to orange, scape elongate, postpedicel brown to black,
>2 times as long as scape, arista yellow; eye bare.

Thorax: Scutum mostly black, mostly yellow pilose, yellow laterally from postpronotum
to postalar callus, sometimes narrowly brown anterior to postalar callus; postpronotum
yellow; postalar callus yellow; scutellum yellow, yellow pilose; pleuron yellow, shining,
bare, brown to black on anterior margin of anepisternum, ventral % katepisternum,
posterior anepimeron and meron; legs mostly yellow, coxae brown, femora sometimes
slightly darker, orange to brown, on basal 7%, tarsi often slightly darker, orange to
brown. Wing infuscated anteriorly in cells bc, c, SC, 4, f2+3 and br anterior to spurious
vein, wing densely microtrichose apically, bare basally, cells br and bm extensively
bare, microtrichose apically, cua bare on basal half, cup bare basally, alula bare;
calypter brown; halter yellow-orange.

Abdomen: Abdomen constricted anteriorly; tergites 1+2 fused, about as wide as
scutellum at base, constricted medially, longer than tergite 4, yellow, small brown spot
on medioanterior edge, large brown marking on medial 14-74; tergites 3 and 4 brown,
posterior edge with broad yellow band, pile on tergites mostly yellow on yellow cuticle
and black on black cuticle except some yellow pile on black cuticle posteriorly on
tergite 4; sternite 1 shining, bare, brown anteriorly, yellow posteriorly; sternite 2 shining,
bare, yellow, often brown medially, usually restricted to lateral edges; sternite 3 yellow
with broad brown to black medial band, mostly black pilose, sometimes yellow pile
medially; sternite 4 broadly brown to black with yellow posterior edge, anterior edge
sometimes narrowly yellow, mostly yellow pile on black cuticle and black pile on yellow
cuticle.

Genitalia: Epandrium longer than tall, mostly covered in short pile; cercus connected to
epandrium posteroventrally, with dorsal lobe, outer ventral corner projecting slightly,
covered in long setae; surstylus pointed, curving inwards, covered in long setae; basal
hypandrium rounded ventrally, apical hypandrium bifurcate, on either side of phallus,
subquadrate with apicodorsal corner projecting further than apicoventral corner,
covered in short pile; phallus laterally compressed, slightly smaller than basal
hypandrium and not projecting much beyond, phallus ends in small point directed
dorsally.

74

Skevington J et al

Female: Differs from male in the following ways: face with yellow pile sometimes
admixed with few black pili, medial facial stripe extending ventrally to oral margin; frons
entirely black pilose; scutellum black or yellow pilose; tergites 1+2 constricted medially
but less so than in male, not as long as in male, yellow anteriorly with brown
anteromedial spot, brown medially, yellow along posterior margin; tergite 5 similar to
tergite 4 of male; sternite 1 almost entirely brown, narrowly yellow posteriorly; sternite 2
yellow with brown medial band and lateral margins brown medially, sparse pile
posteriorly; sternites 3-5 brown with yellow posterior margin, black pilose with some
yellow pili admixed.

See Fig. 10.

Diagnosis

Yellow vertex and black ocellar triangle; face yellow, occasionally with darkened area
below antennae extending to black, narrow medial stripe not reaching the oral margin;
tergites 1+2 fused, longer than tergite 4.

Etymology

Named after the collector of the first Known specimens, P.W. Fattig. Fattig was the
Curator of the Museum in Emory University from 1926 to 1953 and noted by Weems
(1953) to have encouraged his first interest in entomology.

Distribution

This species is known from the south-eastern United States including Florida, Georgia,
Louisiana, Mississippi, Tennessee, Texas and Virginia.

Ecology

This fly is rarely collected. It has been found from early June to late August, with three
specimens having been collected in a Malaise trap sometime from October to
December in Georgia. This species is known from pine, pine-oak, pine-hardwood or
hardwood forests.

Taxon discussion

Mixogaster fattigi differs from M. breviventris Kahl 1897 and M. johnsoni Hull 1941 by
the entirely yellow vertex, entirely yellow metafemur (not dark on apicomedial ’/3) and
yellow lateral margin of 2nd tergite (not dark on posterior 2/3); from M. cubensis Curran
1932 by the uniformly dark postpedicel which is not strongly constricted medially.

New Syrphidae (Diptera) of North-eastern North America 13)

Figure 10.

Mixogaster fattigi, Sp. nov.

a: Dorsal habitus, CNC_Diptera92831 EES]

b: Lateral habitus, CNC_Diptera92831 EES

c: Head, oblique, CNC_Diptera92831 EES]

d: Male epandrium, cercus and surstylus, lateral, CNC1129651; scale bar 200 um ERS]
e: Male genitalia, ventral, CNC1129651; scale bar 200 um | doi|

f: Male hypandrium and phallus, lateral, CNC1129651; scale bar 200 um EES

This species has sometimes been attributed to Howard Weems as he provided a
description of it in his PhD thesis (Weems 1953). However, as Weems himself did not

76

Skevington J et al

consider his thesis a publication in the meaning of the Code (International Commission
on Zoological Nomenclature 1999), the name has remained a nomen nudum. In the
files of the Systematic Entomology Laboratory at the Smithsonian, Chris Thompson
found a complete manuscript by his predecessor, Charles T. Greene, written long
before the Weems' thesis. Despite having not seeing Greene's work, we have
honoured his contribution by including him as an author of the species.

Asingle specimen was sequenced (Table 2).

Common Name

The common name given to the species by Skevington et al. (2019) is Fattig's Ant Fly.

Neoascia guttata Skevington and Moran, sp. n.

ZooBank urn:lsid:zoobank.org:act:A1BD8821-B02E-492C-B2C7-E24209387F7D

Nomenclature

Neoascia undescribed species 17-1 Skevington et al. (2019), page 236

Materials

Holotype:

a. scientificName: Neoascia guttata Skevington and Moran; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Spotted Fen Fly; genus:
Neoascia; subgenus: Neoasciella; specificEpithet: guttata; scientificNameAuthorship:
Skevington and Moran 2019; country: Canada; stateProvince: Nova Scotia; locality:
Lawrencetown, Halifax County; decimalLatitude: 44.644175; decimalLongitude:
-63.344633; samplingProtocol: hand collected; eventDate: 1967-07-19/20; year: 1967;
month: 7; day: 19; verbatimEventDate: 19-20.vii.1967; individualCount: 1; sex: male;
lifeStage: adult; catalogNumber: CNC_Diptera170046; recordedBy: D.M. Wood;
identifiedBy: J.H. Skevington; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

Paratypes:

a. scientificName: Neoascia guttata Skevington and Moran; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Spotted Fen Fly; genus:
Neoascia; subgenus: Neoasciella; specificEpithet: guttata; scientificNameAuthorship:
Skevington and Moran 2019; country: Canada; stateProvince: Alberta; locality:
Jumping Pound Creek, 20 miles West of Calgary; decimalLatitude: 51.040727;
decimalLongitude: -114.753223; samplingProtocol: hand collected; eventDate:
1962-06-23; year: 1962; month: 6; day: 23; verbatimEventDate: 23.vi.1962;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera169737;
recordedBy: W.R.M. Mason; identifiedBy: K. Moran; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

b. scientificName: Neoascia guttata Skevington and Moran; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;

New Syrphidae (Diptera) of North-eastern North America 77

family: Syrphidae; taxonRank: species; vernacularName: Spotted Fen Fly; genus:
Neoascia; subgenus: Neoasciella; specificEpithet: guttata; scientificNameAuthorship:
Skevington and Moran 2019; country: Canada; stateProvince: Alberta; locality: Banff;
decimalLatitude: 51.18; decimalLongitude: -115.57; samplingProtocol: hand collected;
eventDate: 1922-06-2; year: 1922; month: 6; day: 2; verbatimEventDate: 2.vi.1922;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: CNC_Diptera169738;
recordedBy: C.B.D. Garrett; identifiedBy: K. Moran; language: en; institutionCode:
CNC; basisOfRecord: PreservedSpecimen

Description
Size: Body length 4.7 to 5.0 mm; wing length 3.8 to 4.4 mm

Male: Head: Black with metallic sheen; face yellowish pollinose and white pilose; gena
sparsely pollinose and pilose posteriorly; frons shiny, rugose, white pilose on ventral 7,
black pilose dorsally; vertex shiny, smooth, pale pilose; occiput sparsely grey pollinose
and black pilose dorsally, becoming densely silvery white pollinose and white pilose
ventrally; antenna dark brown except orange basoventral 3 of postpedicel; postpedicel
short, about as long as wide; arista slightly longer than postpedicel.

Thorax: Black with metallic sheen; postpronotum shiny, yellowish-white pilose;
mesonotum shiny, yellowish-white pilose; scutellum shiny, yellowish-white pilose;
pleuron greyish-white pollinose except shiny on most of katepisternum, meron and
katatergite; katepisternum with a few white pili dorsally; meron bare; katatergite white
pilose; postmetacoxal bridge absent; coxae dark brown, greyish-white pollinose, white
and black pilose; trochanters orange; pro- and mesofemora narrowly orange basally
and apically, black elsewhere, white pilose; pro- and mesotibiae and tarsi similar with a
bit more orange basally; metafemur orange on basal ’%s and narrowly on apex, black
elsewhere, black pilose; metatibia orange on basal 3 and on apical tip, black medially,
white pilose; tarsi bicoloured, pale pilose; basitarsomere brown except orange apically;
2nd and 3rd tarsomeres orange; apical 2 tarsomeres brown; calypter white; halter
white; wing lightly infuscate, completely microtrichose.

Abdomen: Black except with paired orange spots proximally on 3rd tergite, with
metallic sheen, white pilose.

Genitalia: Male terminalia black pilose; epandrium compact, about same length as
surstylus; cercus elongate, protruding; surstylus broad and elongate, with a few large
bristles on the dorsal proximal corner; distiphallus simple, bag-like; hypandrium
elongate; postgonite simple; phallapodeme straight; ejaculatory apodeme tiny, parasol-
shaped.

Female: Similar to male except for normal sexual dimorphism and the following:
abdomen completely black (Spots absent); metatarsi entirely black.

See Fig. 11.

78

Skevington J et al

Figure 11.
Neoascia guttata, sp. nov.

a: Dorsal habitus, female, CNC_Diptera169737 EES

b: Lateral habitus, female, CNC_Diptera169737 ETS]

c: Lateral head, female, CNC_Diptera169737 EES

d: Lateral habitus, male, Holotype, CNC_Diptera170046 ER
e: Dorsal abdomen, male, Holotype, CNC_Diptera170046 EES
f: Lateral genitalia, male, Holotype, CNC_Diptera170046 EES]

New Syrphidae (Diptera) of North-eastern North America 79

Diagnosis

Postmetacoxal bridge absent. Face long and straight, different from shorter, concave
face found in similar species. Paired spots on tergite 4 of male. Male cerci protruding;
surstyli broad and elongate.

Etymology

This name is from the Latin guttata, meaning dappled, speckled, spotted.

Distribution

This species is known from a single male collected on 19-20 July 1967 in
Lawrencetown, Nova Scotia by Monty Wood and two females from Alberta.

Ecology

This rare fly has been collected from early June to mid-July.

Taxon discussion

The unique face shape (long and straight) suggests that these females are correctly
associated with the males. DNA evidence (Table 2) corroborates this. As the abdominal
pattern varies in Neoascia, it is possible that further dissections will turn up more
specimens of this species that are masquerading as other species. DNA barcodes
suggest that this species is closely related to the North African species N. clausseni
Hauser and Kassebeer 1998. The shape of the face is identical in these two species,
but the abdominal pattern and male genitalia are different.

Common Name

The common name given to the species by Skevington et al. (2019) is Spotted Fen Fly.

Orthonevra feei Moran and Skevington, sp. n.

e ZooBank urn:lsid:zoobank.org:act:66A24081-4FF8-421E-AD79-89E139B6428F

Nomenclature
Orthonevra undescribed species 1 Skevington et al. (2019), page 252

Materials

Holotype:
a. scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:

80

Skevington J et al

Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1987-06-05; year: 1987; month: 6; day: 5; verbatimEventDate: 5.vi.1987;
habitat: Cornus stolonifera; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: Jeff_Skevington_Specimen44217; recordedBy: F.D. Fee; identifiedBy:
K. Moran; language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen

Paratypes:

a.

scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1987-06-05; year: 1987; month: 6; day: 5; verbatimEventDate: 5.vi.1987;
habitat: Cornus stolonifera; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: Jeff_Skevington_Specimen44216; recordedBy: F.D. Fee; identifiedBy:
K. Moran; language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1987-06-05; year: 1987; month: 6; day: 5; verbatimEventDate: 5.vi.1987;
habitat: Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44218; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomencilaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1987-06-07; year: 1987; month: 6; day: 7; verbatimEventDate: 7.vi.1987;
habitat: Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44219; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-13; year: 1990; month: 6; day: 13; verbatimEventDate: 13.vi.1990;
habitat: Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44220; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

New Syrphidae (Diptera) of North-eastern North America

scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;

family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:

Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;

81

eventDate: 1990-06-13; year: 1990; month: 6; day: 13; verbatimEventDate: 13.vi.1990;

habitat: Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44221; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;

family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:

Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;

eventDate: 1990-06-13; year: 1990; month: 6; day: 13; verbatimEventDate: 13.vi.1990;

habitat: Taraxacum; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44222: recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;

family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:

Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;

eventDate: 1990-06-13; year: 1990; month: 6; day: 13; verbatimEventDate: 13.vi.1990;

habitat: Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44223; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: USNM; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;

family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:

Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;

eventDate: 1990-06-13; year: 1990; month: 6; day: 13; verbatimEventDate: 13.vi.1990;

habitat: Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44224:; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;

family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:

Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;

eventDate: 1990-06-13; year: 1990; month: 6; day: 13; verbatimEventDate: 13.vi.1990;

habitat: Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44225; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen

Skevington J et al

scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990;
habitat: Stellaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44226; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990;
habitat: Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44227; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: USNM; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington;
country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott Bog,
Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174; eventDate:
1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990; habitat:
Fragaria; individualCount: 1; sex: male; lifeStage: adult; catalogNumber: Moran and
Skevington 2019; recordedBy: F.D. Fee; identifiedBy: K. Moran; language: en;
institutionCode: ANSP; basisOfRecord: PreservedSpecimen

scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990;
habitat: in cop on flower Fragaria, female feeding on flower; individualCount: 2; sex:
male+female; lifeStage: adult; catalogNumber: Jeff_Skevington_Specimen44229;
recordedBy: F.D. Fee; identifiedBy: K. Moran; language: en; institutionCode: ANSP;
basisOfRecord: PreservedSpecimen

scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1987-06-05; year: 1987; month: 6; day: 5; verbatimEventDate: 5.vi.1987;
habitat: Cornus stolonifera; individualCount: 1; sex: female; lifeStage: adult;

New Syrphidae (Diptera) of North-eastern North America 83

catalogNumber: Jeff_Skevington_Specimen44230; recordedBy: F.D. Fee; identifiedBy:
K. Moran; language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1987-06-06; year: 1987; month: 6; day: 6; verbatimEventDate: 6.vi.1987;
habitat: Cornus stolonifera; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: Jeff_Skevington_Specimen44231; recordedBy: F.D. Fee; identifiedBy:
K. Moran; language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-13; year: 1990; month: 6; day: 13; verbatimEventDate: 13.vi.1990;
habitat: Fragaria; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44232:; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990;
habitat: Fragaria; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44233; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenclaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990;
habitat: Fragaria; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44234; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: USNM; basisOfRecord: PreservedSpecimen
scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990;
habitat: Fragaria; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:

84

Skevington J et al

Jeff_Skevington_Specimen44235; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen

t. scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990;
habitat: Fragaria; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44236; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

u. scientificName: Orthonevra feei Moran and Skevington; nomenciaturalStatus: new
species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Fee's Mucksucker; genus:
Orthonevra; specificEpithet: feel; scientificNameAuthorship: Moran and Skevington
2019; country: U.S.A.; stateProvince: New Hampshire; locality: Coos County, Scott
Bog, Connecticut Lakes; decimalLatitude: 45.217; decimalLongitude: -71.174;
eventDate: 1990-06-14; year: 1990; month: 6; day: 14; verbatimEventDate: 14.vi.1990;
habitat: Fragaria; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen44237; recordedBy: F.D. Fee; identifiedBy: K. Moran;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen

Description
Size: Body length 5.4 to 6.5 mm; wing length 4.1 to 4.9 mm

Description: Head: Black; face concave with lower face projecting forwards along the
oral margin, pale pilose along oral margin but bare above and shiny except with
triangular silver area of pollinosity between antenna and margin of eye; gena pale
pilose, shiny; frontal triangle shiny, pale pilose; vertex black, bare except ocellar
triangle black pilose; occiput pale pilose with some short black setae on dorsal %4 and
shiny except for some white pollinosity along lower eye margin; antenna brown or
brown and orange, scape and pedicel with short, black pile, pedicel about as long as
scape, postpedicel elongate, about three times as long as wide, with minute sensory pit
on inner surface, arista bare; eye bare, with no coloured markings and broadly holoptic.

Thorax: Mesonotum dominantly metallic blue, sometimes with a mix of green, bronze
and or purple colouration, with differently coloured dorsomedial stripes and pale pilose;
scutellum metallic blue, pale pilose; subscutellar fringe absent; postpronotum metallic
blue, pale pilose; postalar callus metallic blue, pale pilose and shiny; pleuron metallic
blue and shiny; anterior anepisternum, katepimeron and meron bare; posterior
anepisternum and anepimeron pale pilose; katepisternum discontinuously pale pilose;
metasternum black and bare; coxa brown, shiny and pale pilose; femora brown except
apex yellow, pale pilose, except metafemur with black setae ventrally and shiny; tibia
brown, except yellow on anterior third and at posterior joint, pale pilose and shiny;
tarsomeres brown except 1st tarsomere yellow and black pilose ventrally; wing entirely
densely microtrichose; halter yellow; calypter white.

New Syrphidae (Diptera) of North-eastern North America 85

Abdomen: Black; tergites 1-4 dull brown medially and shining laterally, black pilose
medially and pale pilose laterally; sternites shining and pale pilose; postabdomen
shining and pale pilose.

Genitalia: Epandrium subquadrate, longer than wide, narrows slightly dorsally; cercus
with laterally compressed, subtriangular sclerotised outer portion covered in long pile
and membranous inner portion; surstylus long, fingerlike, but curving inwards and with
rounded apex, with conspicuous long pile on outer surface and inner surface;
subepandrial sclerite broad, with narrow membranous v-shaped medial section;
hypandrium with patch of macrosetae posterior to postgonite, ejaculatory apodeme
conical; phallapodeme short and straight; postgonite long antler-like structure
projecting ventrally, terminating in three spines; distiphallus elongated apically, narrow,
projecting upwards between surstyli, with sharp pointed apex and long ventral tooth
curving anteriorly ; phallus simple tube within hypandrium.

Female: Same as male except for usual sexual dimorphism.

See Fig. 12.

Diagnosis

Lacks eye markings typical of most Orthonevra species. Legs bi-coloured and
postpedicel longer than scape plus pedicel combined.

Etymology

This species is named after Frank Fee, who collected all known specimens of the
species. Frank was a prolific syrphid collector, returning to the same sites year after
year in an effort to slowly build series around rare or new species.

Distribution

All known specimens were collected from Scott Bog in New Hampshire.

Ecology

All specimens of this rare species were collected in early to mid-June at a single
location. Specimens were collected on Cornus sericea , Fragaria and Taraxacum.

Taxon discussion

Attempts to sequence this species failed. It is clearly closely related to the other
Orthonevra species that lack eye markings. Orthonevra robusta also lacks eye
markings but has entirely black legs and postpedicel shorter than scape plus pedicel
combined.

86

Skevington J et al

Figure 12.

Orthonevra feei, sp. nov.

a: Dorsal habitus, male, Jeff_Skevington_Specimen44221

b: Lateral habitus, male, Jeff_Skevington_Specimen44221

c: Face, oblique, male, Jeff_Skevington_Specimen44221

d: Lateral male genitalia, Jeff_Skevington_Specimen44222, Scale bar 500 um
e: Ventral male genitalia, Jeff_Skevington_Specimen44222, Scale bar 500 um

New Syrphidae (Diptera) of North-eastern North America 87

Common Name

The common name given to the species by Skevington et al. (2019) is Fee's
Mucksucker.

Psilota klymkoi Locke, Young and Skevington, sp. n.

ZooBank urn:lsid:zoobank.org:act:EFDCD11A-7CEF-4364-A960-BFE625547300

Nomenclature

Psilota undescribed species 17-1 Skevington et al. (2019), page 288

Materials

Holotype:

a.

scientificName: Psilota klymkoi Locke, Young and Skevington; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Black Haireye; genus: Psilota
; specificEpithet: klymkol; scientificNameAuthorship: Locke, Young and Skevington
2019; country: Canada; stateProvince: New Brunswick; locality: Albert Co., Caledonia
Grg PNA, Tingley Road; decimalLatitude: 45.8304; decimalLongitude: -64.7784;
samplingProtocol: hand collected; eventDate: 2013-06-10; year: 2013; month: 6; day:
10; verbatimEventDate: 10.vi.2013; habitat: mixed woods at edge of clearcut;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen30434; recordedBy: J. Klymko, S.L. Robinson;
otherCatalogNumbers: JK053333; identifiedBy: M.M. Locke; language: en;
institutionCode: CNC; basisOfRecord: PreservedSpecimen

Paratypes:

a.

scientificName: Psilota klymkoi Locke, Young and Skevington; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Black Haireye; genus: Psilota
; SpecificEpithet: klymkor; scientificNameAuthorship: Locke, Young and Skevington
2019; country: Canada; stateProvince: Quebec; locality: Vaudrieul Co., Summit of
Mount Rigaud; minimumElevationInMeters: 220; decimalLatitude: 45.466389;
decimalLongitude: -74.326389; eventDate: 1995-05-31; year: 1995; month: 5; day: 31;
verbatimEventDate: 31.v.1995; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: CNC_Diptera106482; recordedBy: D.M.Wood; identifiedBy: M.M.
Locke; language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Psilota klymkoi Locke, Young and Skevington; nomencliaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Black Haireye; genus: Psilota
; specificEpithet: klymkol; scientificNameAuthorship: Locke, Young and Skevington
2019; country: U.S.A.; stateProvince: Pennsylvania; locality: Huntingdon County, Ridge
Road, Herod Run; decimalLatitude: 40.636043; decimalLongitude: -77.86749;
eventDate: 1997-06-20; year: 1997; month: 6; day: 20; verbatimEventDate: 20.vi.1997;
habitat: Pastinaca; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen45312; recordedBy: F.D. Fee; identifiedBy: M.M. Locke;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen

88

Skevington J et al

C: scientificName: Psilota klymkoi Locke, Young and Skevington; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Black Haireye; genus: Psilota
; SpecificEpithet: klymkor; scientificNameAuthorship: Locke, Young and Skevington
2019; country: U.S.A.; stateProvince: Pennsylvania; locality: Huntingdon County, Ridge
Road, Herod Run; decimalLatitude: 40.636043; decimalLongitude: -77.86749;
eventDate: 1997-06-20; year: 1997; month: 6; day: 20; verbatimEventDate: 20.vi.1997;
habitat: Pastinaca; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
Jeff_Skevington_Specimen45313; recordedBy: F.D. Fee; identifiedBy: M.M. Locke;
language: en; institutionCode: ANSP; basisOfRecord: PreservedSpecimen

d. scientificName: Psilota klymkoi Locke, Young and Skevington; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Black Haireye; genus: Psilota
; SpecificEpithet: klymkor; scientificNameAuthorship: Locke, Young and Skevington
2019; country: U.S.A.; stateProvince: Pennsylvania; locality: Centre County, Black
Moshannon State Park; decimalLatitude: 40.915503; decimalLongitude: -78.05934;
eventDate: 1999-05-20; year: 1999; month: 5; day: 20; verbatimEventDate: 20.v.1999;
habitat: Pine/blueberry area, on twig; individualCount: 1; sex: female; lifeStage: adult;
catalogNumber: Jeff_Skevington_Specimen45315; recordedBy: F.D. Fee; identifiedBy:
M.M. Locke; language: en; institutionCode: ANSP; basisOfRecord:
PreservedSpecimen

Description
Size: Body length 5.7 to 7.0 mm; wing length 5.1 to 6.0 mm

Male: Head: Face mostly shiny black with scattered pollinosity along central midline of
face and dense pollinosity adjacent to compound eye, mostly black pilose with a few
white pili near the oral margin; frons and vertex black pilose; antenna almost entirely
black, very slightly orangish-brown on ventral side, postpedicel 1.5 times as long as
wide; eye densely yellow pilose.

Thorax: Scutum and scutellum shiny black, entirely black pilose; pleuron shiny black,
pollen-free on anepisternum, katepisternum and anepimeron, other areas of pleuron
with scattered pollinosity, pile almost entirely black, with pale pile only on ventral half of
katepisternum; legs mostly black, with apices of pro- and meso femora, bases of pro-
and meso tibia and ventral side of basitarsomeres dull orange; wing hyaline, with a
small bare area at the base of cell c, anterobasal halves of cells bm and cua bare; alula
completely microtrichose; halter orange; calypter pale brown, pile at edge of calypter
brown.

Abdomen: Tergites black; tergite 1 yellowish-white pilose; tergite 2 with some white
pile anteriorly, otherwise entirely black pilose; tergite 3 almost entirely black pilose, with
a few scattered white pili; tergite 4 with mixed black and white pile, with posterior half of
tergite entirely white pilose; sternites shining, white pilose.

New Syrphidae (Diptera) of North-eastern North America 89

Genitalia: Epandrium compact, about as long as wide; cercus subquadrate, with apex
rounded and slightly wider at base; outer lobe of surstylus inserted at dorsal edge of of
inner lobe, curved downwards smoothly over its entire length and narrowing slightly
towards apex; inner lobe of surstylus broadening very slightly over basal 34, expanding
more abruptly to a rounded tip at apex; hypandrium narrowing smoothly towards apex;
phallapodeme smooth, with ventral projection; postgonite thin, fused to hypandrium,
with tip slightly expanded; phallus with a series of small, ventral spines.

Female: Similar to male, differing as follows: antenna more extensively orange, with
only dorsal edge brown; face with pollen restricted to lateral edges, entirely free of
pollen medially, pile entirely white; frons with mixed black and white pilosity; leg with
Orange areas more extensively orange, tarsomeres entirely orange ventrally;
katepisternum with mixed black and white pilosity on dorsal half as well as ventral;
calypter pale yellowish-white; tergite 2 with pile variable, ranging from entirely black
throughout to white pilose anteriorly, with white pile not reaching lateral edge of tergite.

See Fig. 13.

Diagnosis

Entirely black, shining species with face and frons black pilose. postpedicel 1.5 times
as long as wide. Scutum entirely black pilose, scutellum entirely black pilose, pleuron
entirely black pilose except for some pale pile on ventral half of katepisternum.
Tarsomeres dark brown. Hind coxa mostly white pilose. Tergite 2 mostly black pilose
laterally. Sternite 1 shining. All sternites entirely white pilose.

Etymology

Named after John Klymko, who collected the holotype and suggested that it may be an
undescribed species. John is a zoologist at the Atlantic Canada Conservation Data
Centre, where he works on syrphids and many other animals.

Distribution

This species is known from six specimens from Quebec, New Brunswick and
Pennsylvania.

Ecology

This rare species has been collected from late May to mid-June. It has been found in
mixed woods at the edge of a clearing, in an area of pines and blueberries, on an open
hilltop in mixed forest and nectaring on Pastinaca.

90 Skevington J et al

Figure 13.
Psilota klymkoi, n. sp.

a: Dorsal habitus, Holotype, Jeff_Skevington_Specimen30434 EES]
b: Lateral habitus, Holotype, Jeff_Skevington_Specimen30434 EES
c: Lateral of male genitalia, Holotype, Jeff_Skevington_Specimen30434; scale bar 200um

d: Ventral of male genitalia, Holotype, Jeff_Skevington_Specimen30434; scale bar 200um

Taxon discussion

One COI barcode sequence of this species was obtained (Table 2).

Common Name

The common name given to the species by Skevington et al. (2019) is Black Haireye.

Trichopsomyia litoralis Vockeroth and Young, sp. n.

° ZooBank urn:lsid:zoobank.org:act:1C301DA0-5649-4222-B05D-99C98BEC858F

Nomenclature

Trichopsomyia litoralis Vockeroth manuscript name, in Thompson (1991)

New Syrphidae (Diptera) of North-eastern North America 91

Trichopsomyia undescribed species 1 Skevington et al. (2019), page 320

Materials

Holotype:

a.

scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: Canada; stateProvince: Prince Edward Island; locality: Brackley
Beach, Can. National Park [Prince Edward Island National Park]; decimalLatitude:
46.431111; decimalLongitude: -63.216111; eventDate: 1940-07-30; year: 1940; month:
7; day: 30; verbatimEventDate: 30.vii.1940; individualCount: 1; sex: male; lifeStage:
adult; catalogNumber: CNC_Diptera246378; recordedBy: J. McDunnough;
identifiedBy: J.R. Vockeroth; language: en; institutionCode: CNC; basisOfRecord:
PreservedSpecimen

Paratypes:

a.

scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: Canada; stateProvince: Nova Scotia; locality: South Harbour
Beach; decimalLatitude: 46.878289; decimalLongitude: -60.429056; eventDate:
1983-07-03; year: 1983; month: 7; day: 3; verbatimEventDate: 3.vii.1983; habitat: sand
beach, Ammophila & Lathyrus; individualCount: 1; lifeStage: adult; catalogNumber:
CNC_Diptera110356; recordedBy: J.R. Vockeroth; identifiedBy: J.R. Vockeroth;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: Canada; stateProvince: Prince Edward Island; locality: Green
Gables, Cavendish Beach; decimalLatitude: 46.5018; decimalLongitude: -63.4204;
eventDate: 1967-07-22; year: 1967; month: 7; day: 22; verbatimEventDate:
22.Vil.1967; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC_Diptera246379; recordedBy: D.M. Wood; identifiedBy: J.R. Vockeroth; language:
en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: Canada; stateProvince: Prince Edward Island; locality: Green
Gables, Cavendish Beach; decimalLatitude: 46.5018; decimalLongitude: -63.4204;
eventDate: 1967-07-22; year: 1967; month: 7; day: 22; verbatimEventDate:
22.Vil.1967; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera246380; recordedBy: D.M. Wood; identifiedBy: J.R. Vockeroth; language:
en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:

92

Skevington J et al

Trichopsomyia; specificEpithet: ltoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: Canada; stateProvince: Prince Edward Island; locality: Green
Gables, Cavendish Beach; decimalLatitude: 46.5018; decimalLongitude: -63.4204;
eventDate: 1967-07-22; year: 1967; month: 7; day: 22; verbatimEventDate:
22.vil.1967; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera246381; recordedBy: D.M. Wood; identifiedBy: J.R. Vockeroth; language:
en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: U.S.A.; stateProvince: New Jersey; locality: Cape May;
decimalLatitude: 38.935112; decimalLongitude: -74.906005; eventDate: 06-03; month:
6; day: 3; verbatimEventDate: 3.vi; individualCount: 1; sex: male; lifeStage: adult;
catalogNumber: CNC_Diptera55044; recordedBy: C.W. Johnson; identifiedBy: J.R.
Vockeroth; language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: Canada; stateProvince: Prince Edward Island; locality: Dalvay
House, Can. Nat. Park; decimalLatitude: 46.414905; decimalLongitude: -63.073145;
eventDate: 1940-07-21; year: 1940; month: 7; day: 21; verbatimEventDate:
21.vii.1940; individualCount: 1; sex: male; lifeStage: adult; catalogNumber:
CNC_Diptera55045; recordedBy: G.S. Walley; identifiedBy: J.R. Vockeroth; !anguage:
en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: ltoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: Canada; stateProvince: Prince Edward Island; locality: Dalvay
House, Can. Nat. Park; decimalLatitude: 46.414905; decimalLongitude: -63.073145;
eventDate: 1940-07-21; year: 1940; month: 7; day: 21; verbatimEventDate:
21.vii.1940; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera55046; recordedBy: G.S. Walley; identifiedBy: J.R. Vockeroth; language:
en; institutionCode: CNC; basisOfRecord: PreservedSpecimen

scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: ltoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: Canada; stateProvince: Prince Edward Island; locality: Dalvay
House, Can. Nat. Park; decimalLatitude: 46.414905; decimalLongitude: -63.073145;
eventDate: 1940-07-21; year: 1940; month: 7; day: 21; verbatimEventDate:
21.vii.1940; individualCount: 1; sex: female; lifeStage: adult; catalogNumber:
CNC_Diptera55047; recordedBy: J. McDunnough; identifiedBy: J.R. Vockeroth;
language: en; institutionCode: CNC; basisOfRecord: PreservedSpecimen
scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and

New Syrphidae (Diptera) of North-eastern North America 93

Young 2019; country: U.S.A.; stateProvince: Massachusetts; locality: Gloucester;
minimumElevationInMeters: 16; decimalLatitude: 42.6; decimalLongitude: -70.633333;
eventDate: 1933-06-11; year: 1933; month: 6; day: 11; verbatimEventDate: 11.vi.1933;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: CNC_Diptera9769;
recordedBy: R. Dow; identifiedBy: J.R. Vockeroth; language: en; institutionCode: CNC;
basisOfRecord: PreservedSpecimen

j. scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenciaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: U.S.A.; stateProvince: Virginia; locality: Accomack County, South
End of Assateague Island; decimalLatitude: 38.087029; decimalLongitude: -75.217788;
eventDate: 1997-09-24; year: 1997; month: 9; day: 24; verbatimEventDate: 24.ix.1997;
individualCount: 1; sex: female; lifeStage: adult; catalogNumber: USNMENT247710;
recordedBy: N.E. Woodley; identifiedBy: J.R. Vockeroth; language: en; institutionCode:
USNM; basisOfRecord: PreservedSpecimen

k. scientificName: Trichopsomyia litoralis Vockeroth and Young; nomenclaturalStatus:
new species; kingdom: Animalia; phylum: Arthropoda; class: Hexapoda; order: Diptera;
family: Syrphidae; taxonRank: species; vernacularName: Coastal Psyllid-killer; genus:
Trichopsomyia; specificEpithet: litoralis; scientificNameAuthorship: Vockeroth and
Young 2019; country: U.S.A.; stateProvince: Virginia; locality: Accomack County, South
End of Assateague Island; decimalLatitude: 38.087029; decimalLongitude: -75.217788;
eventDate: 1997-09-24; year: 1997; month: 9; day: 24; verbatimEventDate: 24.ix.1997;
individualCount: 1; sex: male; lifeStage: adult; catalogNumber: USNMENT247711;
recordedBy: N.E. Woodley; identifiedBy: J.R. Vockeroth; language: en; institutionCode:
USNM; basisOfRecord: PreservedSpecimen

Description
Size: Body length 5.5 to 6.0 mm; wing length 4.2 to 4.4 mm

Male: Head: Black; face and frontal triangle white pilose, white microtrichose only on
lateral area touching compound eye; gena shining and white pilose; frontal lunule
brown; vertical triangle shiny, white pilose; occiput silvery white pollinose, white pilose;
scape brown, pedicel brown with orange at apex, black pilose; postpedicel brown, with
a ventral orange stripe, arista bare, orange at base and brown on apical %,
approximately as long as postpedicel. Eye densely covered in short brown pilosity, very
narrowly dichoptic dorsally, separated by about half the width of an ocellus posteriorly
and nearly touching for the length of one ocelli anteriorly.

Thorax: Black, shining, white pilose; proepimeron and anterior anepisternum thinly
white pollinose; scutum shiny, long white pilose; scutellum shiny, long white pilose;
subscutellar fringe short, sparse, white, absent in middle ’% of scutellum; calypter
white; halter whitish-yellow; metathoracic spiracular fringe white. Wing: hyaline,
microtrichose except bare as follows: cell h, basal % cell c, cell br except for scattered
microtrichia, basal % cell r;, basoventral 7 cell bm, base of cell r4,5 and anterobasal
edge of cell cua; alula microtrichose. Legs: coxae black, procoxa silvery white
pollinose, meso- and metacoxa shiny, all coxa white pilose; femora black except apical

94

Skevington J et al

¥~ of pro- and mesofemora yellow and extreme apex of metafemur yellow, white pilose;
protibia yellow, with antero- and ventrolateral 3% brown, mesotibia yellow with anterior 1
brown excluding apex, metatibia brown with basal % yellow, all tibiae white pilose
except apicoventrally on metatibia, which has appressed brownish-orange pile;
protarsus with basitarsomere orange and apical 4 tarsomeres brown, apical 3
tarsomeres black pilose dorsally, tarsus otherwise white pilose; mesotarsomere with
basal 2 tarsomeres orange and apical 3 tarsomeres brown, tarsomere 2 with a few
black pili dorsally, apical 3 tarsomeres black pilose dorsally, tarsus otherwise white
pilose; metatarsus mostly brown, with only apex of basitarsomere, tarsomere 2 and
tarsomere 3 orange, basitarsomere and tarsomere 2 with long white anterodorsal pile,
tarsus otherwise with short white pile, with a few long dorsal black pili on tarsomere 2
and with tarsomeres 3-5 with dorsal pile black.

Figure 14.
Trichopsomyia litoralis n. sp.
a: dorsal habitus, male, CNC_Diptera9769 EES

b: lateral habitus, male, CNC_Diptera9769 EES]
c: face, male, CNC_Diptera9769 ETS]

Abdomen: Black, white pilose; all tergites shining green and white pilose; sternite 1
brown anteriorly and white posteriorly, shiny, white pilose; sternites 2 and 3 brown,
shiny, white pilose, sternite 4 brown, with ventral edge raised smoothly, forming a cup

New Syrphidae (Diptera) of North-eastern North America 95

where the male genitalia sits partially inside, medially thinly white pilose, white
pollinose; 8th segment black, shiny, white pilose.

Genitalia: Epandrium elongate, about as twice as long as tall at the broadest point;
cercus low, triangular, approximately half as tall as broad; surstylus curved, double-
lobed, with large lateral lobe narrowing towards apex, with two broad, dorsal teeth on
inner surface near base and a very small, branching, comb-like ventral lobe near base;
subepandrial sclerite projecting anteriorly between surstyli and broadened at apex;
distiphallus simple, rounded and microtrichose; postgonite broad, approximately twice
as high as long, with a series of small teeth at apex.

Female: Similar to male except for normal sexual dimorphism and: front shiny except
for two lateral pollinose markings on medial 3, white pilose; cell r4,5 more extensively
bare, approximately basal 5 bare; probasitarsomere brown.

See Fig. 14 & Fig. 15.

0.5 mm 0.5 mm

0.2mm

a tas
Figure 15.
Trichopsomyia litoralis n. sp. male genitalia

a: Lateral of genitalia, CNC_Diptera55045 EE

b: Dorsal of genitalia, ses: subepandrial sclerite, CNC_Diptera55045 ERS]

c: Ventral view of surstylus, Il: lateral lobe, vl: ventral lobe, CNC_Diptera55045 EES
d: Phallus, CNC_Diptera55045 EES]

96 Skevington J et al

Diagnosis

Males with eyes separated by about half the width of the anterior ocellus, nearly in
contact for only a short distance; metatarsus with white anterodorsal pile. Female with
cell dm microtrichose at base, sometimes with small bare area along anterior and
posterior margins; scutellar pile long. Both sexes with cell r4.5 bare at base.

Etymology

The word /itoralis is Latin, meaning of the seashore.

Distribution

This species is known from Nova Scotia, Prince Edward Island, Massachusetts, New
Jersey and Virginia.

Ecology

This rare fly has been collected from early June to late September along the east coast
on beaches; one was described as a sand beach with Ammophila and Lathyrus plants
present.

Taxon discussion

A partial COI barcode was obtained from one specimen (Table 2).

Common Name

The common name given to the species by Skevington et al. (2019) is Coastal Psyllid-
killer.

Acknowledgements

Scott Kelso did most of the molecular work that supported the paper. Julie-Anne Dorval
photographed the Anasimyia specimens and edited the Anasimyia genitalia photos. Owen
Lonsdale discussed the homology of the genitalia with the authors. Agriculture and Agri-
Food Canada provided funding for the project. Bill Crins, John Klymko, Gil Miranda and
Menno Reemer reviewed the manuscript and provided useful Suggestions to improve it.

New Syrphidae (Diptera) of North-eastern North America oF

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